Hematopoietic stem/progenitor cells (HSPCs) can self-renew and differentiate into all
hematopoietic lineages, and are extremely sensitive to oxidative stresses. Exposure of HSPCs
to ionizing radiation causes a marked suppression of mature blood cell production in a dose-
dependent manner. However, little information about the long-term effects of low-dose
X-irradiation on the stemness of human HSPCs has been reported. The present study investigated
the biological characteristics of the differentiation and proliferation of low-dose X-irradiated human
CD34
+
HSPCs. Highly purified CD34
+
cells exposed to low-dose X-rays were cultured in liquid and
semisolid media supplemented with an optimal cytokine combination. The liquid medium was
cultured for 14 days; no significant differences in total cell number were observed between non-
irradiated and X-irradiated cultures. The expression levels of cell surface antigens such as CD34,
CD38, and CD45 on cells harvested from X-irradiated culture were similar to those of cells from
non-irradiated culture. However, a significant reduction was observed in the number of burst-
forming unit erythroid (BFU-E) cells between non-irradiated and 0.5 Gy X-irradiated cells in the
myeloid progenitor assay on day 14; moreover, concurrently, radiation-induced γ-H2AX foci were
significantly greater on day 14 than on day 0. These results suggest that low-dose X-irradiation is
associated with clonal growth suppression of BFU-E cells.
Key words: hematopoietic stem/progenitor cells, low-dose X-irradiation, radiosensitivity, burst-
forming unit erythroid cells
Radiation Emergency Medicine 2013 Vol. 2 No. 2 56-62
Proliferation and Differentiation of Human Hematopoietic
Stem/Progenitor Cells Exposed to Low-dose X-irradiation
Masaru Yamaguchi, Satoko Ebina and Ikuo Kashiwakura *
Department of Radiological Life Sciences, Division of Medical Life Sciences, Hirosaki University Graduate School of Health Sciences,
66-1 Hon-cho, Hirosaki, Aomori 036-8564, Japan.
Received 23 February 2013; accepted 29 March 2013
*Ikuo Kashiwakura, Ph.D.: Department of Radiological Life Sciences
Hirosaki University Graduate School of Health Sciences
66-1 Hon-cho, Hirosaki 036-8564, Japan
E-mail: ikashi@cc.hirosaki-u.ac.jp
Copyright © 2013 by Hirosaki University. All rights reserved.
Regular Article
1. Introduction
Ionizing radiation with low-linear energy transfer
such as X-rays generates reactive oxygen species (ROS)
by indirect action of energy on water molecules and
simultaneously generating double-strand breaks (DSBs),
which are known to be lethal, by direct action on DNA
itself, with sequence apoptosis or stress-related responses.
The biological effects of low-dose irradiation have been
investigated, and the cumulative evidence suggests that
the cellular response induced by low-dose irradiation may
differ from that induced by higher doses.
1-4)
Radiation-
induced bystander effects, radioadaptive response, and
activation of the defense system or immune response
represent phenomena that have an important impact
on novel biological responses induced by low-dose
irradiation.
Hematopoietic stem/progenitor cells (HSPCs) can
differentiate into all hematopoietic lineages throughout
the lifetime of an organism and self-renew to maintain