Copyright © 2016 IJRAS, All right reserved
168
International Journal of Research in Agricultural Sciences
Volume 3, Issue 3, ISSN (Online): 2348 – 3997
Study of Genetic Diversity of Berseem Accessions
(Trifolium Alexandrinum L. SPP) With RAPD Markers
S. ROUZ*
Ecole Supérieure d’Agriculture de
Mograne, 1121-El Magren, Tunisie
F. BEN JEDDI
Ecole Supérieure d’Agriculture de
Mograne, 1121-El Magren, Tunisie
M. ZOUAGHI
Institut National Agronomique de
Tunisie, 43 Avenue Charles Nicolle 1082
Tunis-Mahrajène, Tunisie
Abstract – This study aimed to screening genetic diversity
and ancestral relationship between Tunisian berseem
accessions. The analysis of the genetic diversity of 29
accessions of berseem kept and recorded in the fodder index of
Tunisian National Institute of Agronomy has been conducted
via RAPD markers. Several prompts have been tested, but 5
only were selected for their reproductive potential and their
high polymorphism rates. A total of 388 reproducible bands
were obtained during PCR-based amplifications. The bands
have a size ranging between 200 and 2000 pb; though, most of
the bands are in the range of 200 to 600 pb. The correlations
between the accessions have been evaluated by means of
analyses according to the UPGMA method. The extent of
dissimilarity between the accessions varies from 16 to 100%,
which represents a high polymorphism level. The dendrogram,
worked out via the Treecon software, reveals seven sets of
accessions at the 60% threshold. T. berytheum, ancestor of
berseem, clearly emerged from the other accessions. It is of
interest for intensive crop-growing systems and may be used as
parent in selection programs. T. meironense is genetically very
close to the T. alexandrinum var serotinum. Three sets are
generated by T. alexandrinum and T. berytheum crosses. The
present study showed a high genetic diversity of studied
berseem accession. Showed results can be used as a pathway
for preservation and productivity improvement under
different agricultural systems.
Keywords – Trifolium Alexandrinum, Berseem, Genetic
Diversity, T. Berytheum.
I. INTRODUCTION
Zohary and Heller, 1984; Lange and Schifino-
Wittmann, 2000; Le Floc’h et al., 2010 reported that
Tunisia is characterized by a great genetic diversity of the
fodder and pastoral legumes of the genus Trifolium,
having 32 species among a total of 237. According to
Morris & Greene, 2001The berseem (Trifolium
alexandrinum L.) is one of the most important crops in the
Mediterranean agricultural system. The geographical area
of the berseem is the Mediterranean Basin, the ranging
from the Middle East to Irak and Central Asia, like
Pakistan and India (Knight, 1985; Sardana and Narwal,
2000; Badr et al., 2008).
The intensive use of a limited number of improved
varieties of berseem reduces the genetic base of this
species and contributes to genetic vulnerability of its
biodiversity. The ex-situ conservation and the evaluation
of genetic resources of berseem has begun since the 1980s
in the Tunisian National Institute of Agronomy (INAT).
The morphological identification of the genetic variability
in this collection through morphological traits poses a
major issue due to possible inter-crossings amongst
species close to berseem and belonging to the subsection
Alexandrina (Putiyevsky et al., 1975). The RAPD
(Random Amplified Polymorphic DNA) had been used by
Bullita (1995) to characterize varieties and populations of
Trifolium. It demonstrated an efficiency to reveal the
genetic polymorphisms between species and populations
within the species. This method can be used for the
improvement of berseem and for the management of its
genetic resources.
In this work we analyze and try to determine 29 entries
of the genetic diversity of Egyptian clover available in the
collection of the INAT by the RAPD method using
primers which already gave successful results for the other
species of clover (Bullita, 1995).
II. MATERIAL & METHODS
In this study, twenty nine entries of berseem are studied
including a commercial cultivar Miscawi, two local
cultivars, two naturalized cultivars and the rest are natural
populations obtained by exchange with diverse research
institutions, recorded in the fodder index of INAT. The
Origin, the lot number, the name of the genotype and the
code are summarized in table 1.
Table 1. Biologic material, number and origin
Species Number Origin
T. alexandrinum L. 02 TUNISIA-(Local)
T. alexandrinum L. 01 USA
T. alexandrinum L. 02 EGYPT
T. alexandrinum L. 24 INAT
These entries of berseem have been cultivated in
2007/08 in the experimental station of the Agricultural
College of Mograne; situated at a height of 156 m, a
longitude of 10°04’ and a latitude of 36°26’, in a
homogeneous clay-limestone soil. The sowing has been
done in the same date, and the morphological and
agronomic parameters have also been determined (height
of plants, number of stems by plant, date of bloom, color
of flowers, number of cutting, yield dry matter for each
cut,…).
Extraction and Quantification of the DNA
To extract and purify the total DNA from berseem
leaves, we adopted the protocol described by Ben Naceur
(1998) which is a combination of three methods of
extractions (Murray and Thomson, 1980; Saghai-Maroof
et al., 1984; Webb and Knapp, 1990) bringing CTAB.
A representative sample of 1g of leaves from every
entry has been taken from 10 plants of berseem and
crushed in the mortar, in the presence of liquid nitrogen,
until obtaining of a very fine powder. Each mix has been
transferred to a 10 ml tube, adding 7.2 ml of extraction