a SciTechnol journal Research Article
Leo Daniel et al., J Plant Physiol Pathol 2013, 1:2
http://dx.doi.org/10.4172/jppp.1000105
Journal of Plant
Physiology & Pathology
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Technology and Medicine
Effect of Polymeric Additives,
Adjuvants, Surfactants on
Survival, Stability and Plant
Growth Promoting Ability of
Liquid Bioinoculants
Leo Daniel Amalraj E
1
*, Venkateswarlu B
1
, Suseelendra Desai
1
,
Praveen Kumar G
1
, Mir Hassan Ahmed SK
1
, Meenakshi T
1
,
Uzma Sultana
1
, Sravani Pinisetty
1
and Lakshmi Narasu M
2
Abstract
The present study was conducted to evaluate the effect of polymeric
additives, adjuvant and surfactants for their ability to support
growth, shelf-life stability and bio-effcacy of liquid bioinoculants
(Bacillus megaterium var. phosphaticum, Azospirilum brasilense
and Azotobacter chrococcum). Liquid inoculants formulated with
2% polyvinylpyrollidone (PVP 30 K), 0.1% carboxy methylcellulose
(CMC-high density) and 0.025% Polysorbate 20 promoted
long-term survival of Bacillus megaterium var. phosphaticum,
Azospirillum and Azotobacter with 5.6 x 10
7
, 1.9x10
8
and 3.5x10
7
cfu ml
-1
, respectively after 480 days of formulation when stored
at 30
o
C. Maize seeds treated with consortium of PSB and
Azospirillum brasilense enhanced plant growth positively by a
multitude of synergistic mechanisms when compared to single
inoculant application.
Keywords
Liquid inoculants; Cell protectants; Shelf life; Growth promotion
Introduction
Bioinoculants contain benefcial bacteria that enhances plant
growth by the virtue of its nutrient solubilisation [1], nitrogen
fxation [2], phytohormone production [3,4] and induction
of defence mechanism [5-7] abilities. In India, Azospirillum,
Azotobacter, Rhizobium, potash mobilizing bacteria, phosphate and
zinc solubilising bacteria were included under Fertilizer control
order, 1985 in view of nourishing the organic agriculture through
the application of quality bioinoculants. Bhattacharyya and Kumar
[8], stated that, bioinoculants manufactured in India are mostly solid
carrier based and have a shelf life of only six months. Tese carrier-
based inoculants are inherent with certain constraints like lower
shelf-life, poor survival under adverse environmental conditions,
high degree of contamination, and inconsistent feld performances.
Tere have been many attempts to fnd alternatives for carrier based
inoculants and, also to enhance viability of microorganisms in the
inoculants. Besides, it requires a signifcant amount of processing,
*Corresponding author: Leo Daniel Amalraj E, Division of Crops Sciences,
Central Research Institute for Dryland Agriculture, Santoshnagar, Hyderabad,
Andhra Pradesh, India-500059, E-mail: leodanielamalraj@hotmail.com
Received: March 02, 2013 Accepted: Apirl 29, 2013 Published: May 03, 2013
such as mining, drying, milling, neutralizing and also becomes
expensive in sterilization, formulation and transportation processes.
Liquid inoculant formulations (LIF) could be a possible
solution to the foresaid problems. It containing not only the desired
microorganisms and their nutrients but also special cell protectants
or additives that promote for longer shelf life and tolerance to
adverse conditions [9]. Moreover, additives in the broth will improve
inoculants quality, such as including better adhesion to seed [10],
stabilizing the product, binding or inactivating soluble seed coat
toxins. Many kinds of polymers have been used for inoculant
production because of their ability to limit heat transfer, their good
rheological properties and high water activities. Tese polymers,
such as sodium alginate [11], gum Arabic [12], polyvinyl pyrrolidone
(PVP) [13], polyvinyl alcohol (PVA) [14] and polyethylene glycol
(PEG) [15] are normally used as adhesive when they are applied
to seed. Polymers are also used to entrap bacteria in microbeads
[16,17]. Tey are also soluble in water and make convenient for seed
application a simple process for farmers. Te selection of ingredients
was based on their properties, such as solubility in water, non-toxicity
and complex chemical nature, which prevents microorganisms in
the soil from rapidly degradation. It is always a herculean task for a
single inoculant to instigate signifcant impact on the plant growth.
Co-inoculation can beneft plant growth by diferent mechanisms
[11]. Hence, we evaluated the optimum concentration of polymers,
surfactants and adjuvants for LIF that could sustain the bacterial shelf
life and also studied its bioefcacy in pot culture using maize (Zea
mays. L) as test crop.
Materials and Methods
Microorganisms
Bacillus megaterium var. phosphaticum (BMPSB 01), Azospirillum
brasilense (ASP 20) and Azotobacter chrococcum (AZB 29) were taken
from microbial culture bank, Central Research Institute for Dryland
Agriculture, Hyderabad, India. Tey were isolated from rhizospheric
soil of maize plants in Hayathnagar Research Farm, Hyderabad, India.
Growth medium for liquid inoculants
A modifed Pikovaskaya’s broth composed of (gL
-1
) yeast extract:
5.0, dextrose: 10.0, Ca
3
(PO
4
)
2
: 5.0, KCl: 0.5, MgSO
4
: 0.2, MnSO
4
: 0.1,
FeSO
4
: traces, (NH
4
)
2
SO
4
: traces, glycerol: 10 ml was used to culture
Bacillus megaterium var phosphaticum. Modifed Jenson’s broth was
used to culture Azotobacter which composed of sucrose: 20, K
2
HPO
4
:
1.0, MgSO
4
: 0.5, Nacl: 0.5, FeSO
4
: 0.1, Na
2
MoO
4
: 0.005, glycerol: 10
ml and for Azospirillum, modifed nitrogen free bromothymol blue
medium containing L-Malic acid: 5.0, K
2
HPO
4
: 0.5, MgSO
4
: 0.2, Nacl:
0.1, CaCl
2
: 0.02, Na
2
MoO
4
: 0.002, MnSO
4
: 0.01, FeEDTA (1.64% w/v):
4.0 ml, glycerol: 10 ml, bromo thymol blue (0.5% w/v in ethanol): 3.0
ml, KOH: 4.5, biotin: 0.1 were used as a basal media to evaluate the
efect of additives on cell growth.
Screening diferent concentration of additives on bacterial
growth
All the three media were amended with diferent concentrations
of additives as follows: PVP at 1.0, 2.0, 3.0 and 5.0% (w/v); PEG at