ICANCER RESEARCH 53.3752-3757. August 15,1993]
Pharmacokinetics and Toxicology of Immunotoxins Administered into the
Subarachnoid Space in Nonhuman Primates and Rodents
Karin Muraszko, Cynthia Sung, Stuart Walbridge, Larry Greenfield, Robert L. Dedrick, Edward H. Oldfield, and
Richard J. Youle
Biochemistry Section, Surgical Neurology Branch, National Inslilule of Neurological Diseases ami Stroke /K. M., S. W., E. H. O., R. J. Y.¡,and Biomedicai Engineering and
Instrumentation Program, Intramural Research Resources, National Center for Research Resources ¡C.S., R. D.], N1H. Bethesda, Maryland 20892, and Cetus Corp., Emeryville,
California "460ft /L. G.J
ABSTRACT
Immunotoxins have been suggested as possible therapeutic agents in
patients with leptomeningeal carcinomatosis. The pharmacokinetics, sta
bility, and tuxicity of immunotoxins injected into the i.t. space were ex
amined in rats and rhesus monkeys. Monoclunal antibodies specific for the
human (454A12 and Jl) and rat (OX26) transferrin receptors were cou
pled to recombinant ricin A chain. In monkeys, the maximally tolerated
dose of the anti-human transferrin receptor immunotoxin (454A12-rRA)
was a dose that yielded a nominal cerebrospinal fluid (CSF) concentration
of approximately 1.2 x IO"7 M. In rats, the 10% lethal dose (LD10) of the
anti-human transferrin receptor immunotoxin was a dose yielding a nom
inal CSF concentration of 8.8 x III M whereas the LDIO of the anti-rat
transferrin receptor immunotoxin (OX26-rRA) was a dose yielding a nom
inal (SI concentration of 1.2 x II) M.Thus, the species-relevant antibody
resulted in toxicity at a concentration one-seventh that of the immunotoxin
with the irrelevant antibody. A comparison of the area under the concen
tration curve at the 1.l>m for rats with the area under the concentration
curve at the maximally tolerated dose in monkeys and humans shows that
the species-relevant immunotoxin was a better predictor of the toxic dose
of the anti-transferrin receptor immunotoxin in humans than the irrele
vant immunotoxin. The pharmacokinetics of the 454A12-rRA immuno
toxin within the CSF of monkeys showed a biphasic clearance with an
early-phase half-life of 1.4 h and a late phase half-life of 10.9 h. The
clearance was 4.4 ml/h or approximately twice the estimated clearance due
to bulk flow of CSF. Loss by degradation was ruled out because iminii-
noblot analysis showed that the immunotoxin was stable for up to 24 h
after administration. Possible losses in addition to sampling include dif
fusion into brain tissue and transcapillary permeation. The apparent vol
ume of distribution was 10.1 ml or approximately three-fourths the total
CSF volume of the monkey. Dose limiting toxicity corresponded with the
selective elimination of Purkinje cells in both rats and monkeys and was
manifested clinically as ataxia and lack of coordination. The onset of
ataxia in monkeys occurred within 5 days and, in the more mild form, was
reversible with time. There was evidence of only minimal inflammation
within the CSF, and there were no signs of systemic toxicity. Immunotox
ins injected into the subarachnoid space may have potential for treatment
of leptomeningeal carcinomatosis.
INTRODUCTION
Meningea! carcinomatosis occurs in 5-20% of all cancer patients,
the majority caused either by breast carcinoma or lung carcinoma (1).
There are recent indications that the incidence of meningea! carcino
matosis is increasing as patients survive longer with improved sys
temic therapy. Yet, in two-thirds of cases, it occurs when systemic
disease is stable or in complete remission (1-7). These patients have
an exceedingly poor prognosis. Patients able to tolerate maximal
therapy (i.t. methotrexate and whole-brain irradiation) have a 6-7-
month mean survival and less than 15% are alive after 1 year, even
with aggressive therapy (7-15). In children, the incidence of CSF1
Received 12/30/92; accepted 6/8/93.
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1The abbreviations used are: CSF, cerebrospinal fluid; MTD. maximally tolerated
dose; LDu>. 10% lethal dose; AUC. area under the concentration curve; CNS. central
spread of primary CNS tumors such as medulloblastomas and ependy-
momas can be quite high. As many as 44% of children with medullo-
blastoma have evidence of spread via CSF pathways, and 31% of
children who die from ependymomas have evidence of CSF métas
tases (16). Malignant gliomas, particularly in the pediatrie population,
may also have CSF spread. Thus, CSF spread of cancer causes sig
nificant morbidity, and the choice of treatments now available is
limited.
Monoclonal antibodies coupled to protein toxins such as ricin,
called immunotoxins, represent a potent and cell type-selective new
class of chemotherapeutic agents (17-19). Immunotoxins can specif
ically kill a variety of tumor cell types in vitro and in vivo, however,
systemic delivery of immunotoxins is hampered by rapid clearance
into nontarget tissues and slow transport into tumor tissue. Regional
delivery may overcome some of these limitations to immunotoxin
delivery, particularly for therapy of CNS cancer, where the blood-
brain barrier additionally impedes macromolecule delivery (8, 9, 20-
22). In animal models of leptomeningeal carcinomatosis, immunotox
ins delivered directly into the CSF prolong survival in tumor-bearing
animals (20, 23). These and other studies have demonstrated a poten
tial therapeutic benefit of administration of immunotoxins to the CSF
in patients with leptomeningeal carcinomatosis (20, 24, 25).
Our study assessed the pharmacokinetics, stability, and toxicity of
immunotoxins in the CSF of rats and rhesus monkeys. To address the
nonspecific toxicity of immunotoxins in animals and to thoroughly
identify the potential of species-specific, antibody-mediated toxicity
in an animal model we constructed and studied a human-specific
immunotoxin as well as an analogous rat-specific immunotoxin.
MATERIALS AND METHODS
The immunotoxins studied were: (a) 454A12-rRA, an anti-human transfer
rin receptor monoclonal antibody of the IgGl isotype (454A12) (2ft) conju
gated to recombinant ricin A chain (rRA); (b) Jl-rRA, an IgG2a monoclonal
antibody to the human transferrin receptor coupled to rRA (26); and (c)
OX26-rRA. an IgG2a monoclonal antibody to the rat transferrin receptor (27)
conjugated tu rRA. Conjugation of rRA to the antibodies was carried out as
described previously (28).
Protein Synthesis Assay
The cytotoxic effects and potency of the anti-rat transferrin receptor immu
notoxin were tested on a rat myeloma cell line, Y3-Agl.2.3 (29) and a rat
bladder carcinoma cell line. NBT II (30). The myeloma cell line was main
tained in RPMI 1640 containing 10% fetal calf serum, 10 HIM4-(2-hydroxy-
ethyl)-l-piperazineethanesulfonic acid, 20 fig/ml gentamicin, and 2 RIM
glutamine; the bladder cancer cell line was maintained in Dulbecco's modified
Eagle's medium with 10% fetal calf serum, 2 ITIMglutamine, 1 IBM sodium
pyruvate, 0.1 IBM nonessential amino acids, and 10 /xg/ml gentamicin. The
protein synthesis rate was assayed as previously described (24).
nervous system; PBS. phosphate-buffered saline; i.t., intrathecal; p.c., percutaneous; TfR,
transferrin receptor; In-DTPA. indium diethylenetriaminepentaacetic acid.
3752
Research.
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