Bioresource Technology41 ( 1992) 59-63
Non-destructive Estimation of Biomass of Salix
dasyclados
Theo Verwijst & Nils-Erik Nordh
The Swedish University of Agricultural Sciences, Department of Ecology and Environmental Research, Section of
Short Rotation Forestry, Box 7072, S-750 07 Uppsala, Sweden
(Received 25 January 1991; accepted 25 June 1991 )
Abstract
Common non-destructive methods for biomass
estimation of willow developed for Salix viminalis
stands perform poorly when applied to Salix
dasyclados, because of differences between these
species in stem shape and branching pattern. A
method has been developed by which the weight of
a single stem, consisting of several shoots with
apical dominance, is related to the sum of the
cross-sectional areas of the shoots. This method
gives a ,good fit of the data with a linear regression
model, and performs better than models which
include stem height in the predictor variable.
Key words: Biomass estimation, Salix dasyclados,
willow.
INTRODUCTION
During the last decade, short rotation forestry
research in Sweden has focused on the use of
willow (Salix spp.)to produce biomass for energy
purposes (Sir6n, 1983). Descriptive methods for
stand characterisation for assessment of biomass
dynamics (Nilsson, 1981, 1982) were developed
simultaneously with programmes for breeding
and selection of willow clones (Sir6n, 1983; Ager
et al., 1986; Gullberg, 1988).
Methods for biomass estimation were
developed using Salix viminalis, but the earlier
breeding programmes also released clones of
other Salix species. Some of these species display
growth forms which differ markedly from the
erect physiognomy of Salix viminalis. An extreme
example of such different growth form is found in
the Salix dasyclados clone 77075, which produces
Bioresource Technology 0960-8524/92/S05.00 © 1992
Great Britain
'bow-shaped' stems, often with an almost horizon-
tally oriented lower stem part. Clone 77075 pro-
duces curved branches, starting at different
heights above the ground. Many of these branches
show apical dominance.
Clone 77075 and some other clones with the
same growth form are currently being tested on a
large scale and an annual biomass estimation is
needed for the evaluation of these trials. Some of
the stands are relatively small and destructive
sampling would influence their future develop-
ment. Furthermore, destructive sampling is time-
consuming, especially in stands which have grown
during more than one season, and therefore a suit-
able non-destructive method is preferable in most
cases.
The non-destructive biomass estimation
methods developed for Salix viminalis require
stem weight to be related to stem diameter. If
stems of Salix dasyclados clone 77075 are har-
vested close to the soil surface while diameter is
measured at a higher level, a series of diameter
scores has to be related to one stem weight.
Relating one weight score with a series of dia-
meter scores can be avoided by measuring the
diameter at harvest level, but this may cause large
errors due to the taper of the stems, which is more
pronounced at the stem base (Nilsson, 1982). The
number of diameters to be measured per har-
vested stem will largely be a function of the height
above the ground at which diameter is measured.
The inclusion of stem height as a predictor var-
iable for weight brings about comparable prob-
lems.
Biomass estimation on Salix dasyclados has
been carried out in Finland (Hyt6nen, 1985;
Hyt6nen et al., 1987), but the clones used in these
studies did not have a branching pattern which
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