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International Journal of Fisheries and Aquatic Studies 2016; 4(4): 125-129
ISSN: 2347-5129
(ICV-Poland) Impact Value: 5.62
(GIF) Impact Factor: 0.352
IJFAS 2016; 4(4): 125-129
© 2016 IJFAS
www.fisheriesjournal.com
Received: 17-05-2016
Accepted: 18-06-2016
Maulilio J Kipanyula
Department of Veterinary
Anatomy, Faculty of Veterinary
Medicine, Sokoine University of
Agriculture, P. O. Box 3016,
Chuo Kikuu, Morogoro Tanzania
Keziah W Maina
Department of Veterinary
Anatomy, Faculty of Veterinary
Medicine, Sokoine University of
Agriculture, P. O. Box 3016,
Chuo Kikuu, Morogoro Tanzania
Correspondence
Maulilio J Kipanyula
Department of Veterinary
Anatomy, Faculty of Veterinary
Medicine, Sokoine University of
Agriculture, P. O. Box 3016,
Chuo Kikuu, Morogoro Tanzania
Morphological and adaptational changes associated
with fish migration from fresh to marine water bodies
Maulilio J Kipanyula and Keziah W Maina
Abstract
Expansion of the aquatic industry and increased preference on white over red meat has resulted into
intensification of aquaculture practices. Inland capture fisheries in Africa are also on the increase while
marine capture has stagnated over the last 20 years. This has led to unconventional fish farming where by
some marine and fresh water fish species are increasingly farmed in fresh and sea waters respectively.
Although some fish species can naturally survive on fresh and sea water environments, the transition
between the two water bodies requires morphological changes to ensure survival and optimal
productivity. Morphological changes take place concurrently, with the physiological adaptations which
are critical for survival in the new environment. Artificial hatcheries can simulate environmental
conditions which favour such transitions, and therefore proper understanding of the underlying
anatomical and physiological changes is crucial. In this paper, we summarize and discuss available
reports on the anatomical and physiological changes associated with migration of fish from fresh to sea
water and vice versa. Understanding of species specific adaptational changes is crucial as farmers engage
in intensive fish farming involving transfer from hypoosmotic to hyperosmotic water environments and
vice versa.
Keywords: Smoltification, fish, fish hatcheries, parr.
Introduction
Some fish species inhabit both freshwater and seawater bodies and routinely migrates between
the two environments
[1]
. The transition between water bodies lead to anatomical changes such
as: change in pigmentation, body shape and length, emergence and growth of teeth on the
maxilla, mandible and tongue; growth and changes in shape of integumentary folds adjacent to
the cloacal opening; growth and change in shape of the auxiliary appendage of the pelvic fin;
and growth in the scales with respect to radius and number of circuli. Additionally,
adaptational morphological changes also occur in the gills, esophagus, kidney and intestines
[1,
6-9]
. Migration between fresh and marine water ecosystems is often associated morphological
changes referred to as smoltification or parr-smolt transformation and the resulting migrant is
termed a smolt, which is characteristic of juvenile fish. In this context there are two groups of
fish namely as; catadromous and anadromous fish
[2]
. Catadromous fish are born in saltwater,
then migrate to freshwater as juveniles where they grow into adults, before migrating back to
the ocean to spawn. The catadromous fish include; true eels (Anguillids spp), thin-lipped grey
mullets (Liza ramada) and European flounders (Platichthys flesis). This metamorphosis
involve alterations in lipid metabolism, osmoregulation, oxygen transport, buoyancy, growth,
colour, shape, rheotaxis and behavior are preparatory events to maximize survival during
downstream migration, ocean entry and long distance feeding migrations in the marine
environment
[1].
In anadromous fish reproduction is limited to rivers which have few predators thus increasing
chances of survival during the early stages. The river ecosystem supports limited growth rate
and size of fish, therefore by leaving at a certain life stage and entering ocean which has rich
marine food supply, they increase their reproductive success. The age at which smoltification
occurs and its intensity differ among species. For example; in pink salmon (Oncorhynchus
gorbuscha) and chum salmon (Oncorhynchus keta) seaward migration takes place soon after
emergence from the gravel and the rest of the life cycle and growth takes place in the sea. The
Charr (Salvelinus spps) are entirely fresh water or spend only a few weeks feeding in sea water
during summer.