Short Communication
Long-term Cryoconservation and Stability of Vitamin C
in Serum Samples of the European Prospective
Investigation into Cancer and Nutrition
Mazda Jenab,
1
Sheila Bingham,
2
Pietro Ferrari,
1
Marlin D. Friesen,
1
Wael K. Al-Delaimy,
1
Robert Luben,
3
Nick Wareham,
4
Kay-Tee Khaw,
5
and Elio Riboli
1
1
Nutrition and Hormones Group, IARC-WHO, Lyon, France;
2
Medical Research Council Dunn Human Nutrition Unit;
3
Department of Community Medicine,
4
Institute of Public Health, and
5
Clinical Gerontology Unit,
University of Cambridge, Cambridge, United Kingdom
Abstract
Plasma vitamin C level may be associated with risk of some
chronic diseases. The rapid degradability of vitamin C in
biological samples necessitates its stabilization with meta-
phosphoric acid or similar agents. However, in most cohort
studies, prospectively collected biological samples are not
treated with stabilizing agents before long-term frozen
storage and it is not known whether vitamin C can be
properly measured in such samples. The objective of this
study was to determine the degree of vitamin C degradation
in plasma samples stored without stabilization for 7 to 11
years at À196°C. Spearman’s correlation coefficients indi-
cate a moderate correlation between baseline and final
plasma vitamin C levels in both men (r = 0.57, P < 0.0001)
and women (r = 0.52, P < 0.0001). Samples were also
categorized based on low or high baseline levels of plasma
vitamin C, with the latter category showing the highest rate
of loss per year of frozen storage in men (1.96 Mmol/L, P
value for difference <0.0001; percent loss 24.6%) and women
(2.35 Mmol/L, P value for difference <0.0001; percent loss
24.2%), as determined by multiple regression analysis
adjusted for smoking status, age, and body mass index. In
men, both baseline and final plasma vitamin C values were
lower in smokers than never smokers, but for both men and
women the rate of vitamin C loss during storage was not
significantly different between smokers and never smokers.
The results of this study show that vitamin C can be
measured with reasonable reliability in plasma samples
frozen for long periods of time without addition of any
stabilizing agents. (Cancer Epidemiol Biomarkers Prev
2005;14(7):1837 – 40)
Introduction
Vitamin C from the diet has been suggested to be inversely
associated with risk of some cancers (1, 2) and cardiovascular
disease (2), likely due to its antioxidant properties (3). With the
advent of appropriate analysis techniques, measures of
vitamin C in biological samples have been used as a biomarker
of health status [e.g., in the study of gastric cancer (4, 5) and
total mortality (6)]. Because vitamin C in biological samples
can easily degrade or be readily oxidized, it is often stabilized
before sample storage by the addition of stabilizing agents
such as EDTA (a metal chelator), perchloric acid, DTT, or
metaphosphoric acid, all of which act as protein-precipitating
agents (7). However, in the case of large, multipurpose
prospective studies, biological samples are collected and often
stored for many years before analysis for a multitude of
biomarkers. Thus, blood samples are not always treated
specifically for the preservation of vitamin C before storage.
Present literature on vitamin C stability deals with losses
during the handling, processing, and freezing procedures of
blood samples (7-12), after short periods of frozen storage
without stabilization (11, 12) or for longer time periods of up to
6 years with stabilization (13, 14). However, there is currently
very little information on the feasibility, stability, and validity
of vitamin C analysis in blood samples stored by freezing at
very low temperatures (À196jC) for very long periods of time
without the addition of stabilizing compounds such as
metaphosphoric acid. Thus, the objective of this study was to
determine the stability of vitamin C in plasma samples stored
for 7 to 11 years in liquid nitrogen and not treated with any
stabilizing agents during storage compared with samples
analyzed at baseline with stabilization. The samples were
collected from a subset of subjects enrolled in the European
Prospective Investigation into Cancer and Nutrition (EPIC)
study.
Materials and Methods
The EPIC Study and the EPIC-Norfolk Subcohort. The
EPIC study (15-17) is an ongoing prospective multicenter
cohort study with 521,468 subjects from 23 centers in 10
European countries. The EPIC biorepositories are the world’s
largest, hosting over 9 million aliquots maintained at À196jC.
For the present study, subjects were selected from the Norfolk
(Cambridge, United Kingdom) center of the EPIC study (EPIC-
Norfolk). The enrollment, recruitment, and sample collection
Cancer Epidemiology, Biomarkers & Prevention 1837
Cancer Epidemiol Biomarkers Prev 2005;14(7). July 2005
Received 1/24/ 05; revised 3/29/ 05; accepted 4/14/ 05.
Grant support: ‘‘Europe Against Cancer’’ Programme of the European Commission; Ligue
contre le Cancer (France); Socie ´te ´ 3M (France); Mutuelle Ge ´ne ´rale de l’Education Nationale;
Institut National de la Sante ´ et de la Recherche Me ´dicale; German Cancer Aid; German Cancer
Research Center; German Federal Ministry of Education and Research; Danish Cancer Society;
Health Research Fund (FIS) of the Spanish Ministry of Health; the participating regional
governments and institutions of Spain; Cancer Research UK; Medical Research Council, UK;
the Stroke Association, UK; British Heart Foundation; Department of Health, UK; Food
Standards Agency, UK; the Wellcome Trust, UK; Greek Ministry of Health; Greek Ministry of
Education; Italian Association for Research on Cancer; Italian National Research Council;
Dutch Ministry of Public Health, Welfare and Sports; Dutch Ministry of Health; Dutch
Prevention Funds; LK Research Funds; Dutch Zorg Onderzoek Nederland; World Cancer
Research Fund; Swedish Cancer Society; Swedish Scientific Council; Regional Government of
Skane, Sweden; and Norwegian Cancer Society.
The costs of publication of this article were defrayed in part by the payment of page charges.
This article must therefore be hereby marked advertisement in accordance with 18 U.S.C.
Section 1734 solely to indicate this fact.
Requests for reprints: Mazda Jenab, Nutrition and Hormones Group, IARC-WHO, Lyon,
France. Phone: 33-4-72-738082; Fax: 11-33-4-72-738361. E-mail: jenab@iarc.fr
Copyright D 2005 American Association for Cancer Research.
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