1 Title: Evaluating species distribution models with discrimination accuracy is 1 uninformative for many applications 2 3 Running title: Evaluating niche models with occurrences 4 5 Authors: Dan L. Warren 1,2,3* , Nicholas J. Matzke 4 , and Teresa L. Iglesias 3,5 6 7 * To whom correspondence should be addressed: dan.l.warren@gmail.com 8 1. Senckenberg Biodiversity and Climate Research Centre, Frankfurt, Germany 9 2. Biodiversity and Biocomplexity Unit, Okinawa Institute of Science and 10 Technology, Okinawa, Japan 11 3. Department of Biology, Macquarie University, Sydney, Australia 12 4. Department of Biological Sciences, University of Auckland, Auckland, New 13 Zealand 14 5. Physics and Biology Unity, Okinawa Institute of Science and Technology 15 16 17 18 Author Contributions: All authors contributed to study design and manuscript 19 preparation. Simulations were coded and run by DLW, literature review was 20 performed by TLI. Post hoc analyses were performed by TLI and DLW. 21 . CC-BY 4.0 International license not certified by peer review) is the author/funder. It is made available under a The copyright holder for this preprint (which was this version posted June 27, 2019. . https://doi.org/10.1101/684399 doi: bioRxiv preprint