Indarctos (Ursidae, Mammalia) from the Spanish Turolian (Upper Miocene) P. Montoya, L. Alcalá & J. Morales Montoya, P., L. Alcalá & J. Morales. Indarctos (Ursidae, Mammalia) from the Spanish Turolian (Upper Miocene). — Scripta Geol., 122: 123-151, 7 figs., 2 pls, Leiden, June 2001. P. Montoya, Departament de Geologia, Universitat de València, Dr Moliner 50, 46100 Burjassot, Spain, e-mail: pmontoya@uv.es; L. Alcalá & J. Morales, Museo Nacional de Ciencias Naturales (CSIC), José Gutiérrez Abascal 2, 28006 Madrid, Spain, e-mail: alcala@mncn.csic.es mcnm166@mncn.csic.es. Key words: Mammalia, Ursidae, Miocene, Spain. Abundant fossils of the bear Indarctos atticus (Weithofer, 1888) from the Lower Turolian (MN11) site of Crevillente 2 (Alicante, Spain) and some new material from the Middle Turolian (MN12) of Con- cud and Valdecebro 5 (Teruel, Spain) are described. On the other hand some dental remains from Crevillente 2 are identified as Indarctos sp. cf. I. vireti Villalta & Crusafont, 1943 on the basis of their small size. The status of different Indarctos species are discussed. Contents Introduction ......................................................................................................................... 123 Systematics ........................................................................................................................... 124 Indarctos atticus (Weithofer, 1888) ............................................................................... 124 Indarctos sp. cf. Indarctos vireti Villalta & Crusafont, 1943....................................... 141 Stratigraphic and geographic distribution of the species of the genus Indarctos ....... 143 Acknowledgements ............................................................................................................. 144 References ............................................................................................................................. 144 Introduction In Spain, Indarctos atticus has been published and described mainly from several localities of the Lower and Middle Turolian (MN11 and MN12) in the Teruel Basin: Concud (Hernández Pacheco, 1930a; Crusafont, 1962; Alcalá, 1994), Los Mansuetos (Adrover, 1963), Valdecebro 5 (Adrover et al., 1986) and Puente Minero (Alcalá et al., 1991; Alcalá, 1994). Outside the Teruel Basin it has only been reported from the Late Vallesian of Terrassa (Pons Moyà, 1989-90) and from Crevillente 2 (Alcalá et al., 1987; Alcalá & Montoya, 1989-90; Montoya, 1994). In this paper we describe the material from Crevillente 2, consisting of a relatively rich sample of dental and postcranial remains, and also some unpublished material from Valdecebro 5 and Concud (Cerro de la Garita). We also study a jaw fragment, and an incomplete lower carnassial, from the lower levels of Crevillente 2, which are clearly smaller than those of I. atticus and, on the other hand, very similar to those of I. vireti; they are assigned with some reserves to that species. The mammals from Crevillente (prov. Alicante, Spain) have been studied in numerous papers, among others Montenat & Crusafont (1970), Montenat (1973), de Bruijn et al. (1975), Alcalá et al. (1987), Alberdi & Montoya (1988), Cerdeño (1989), Alcalá & Montoya (1989-90), Freudenthal et al. (1991, 1998), Montoya & Morales (1991), van der Made et al. (1992), Martín Suárez & Freudenthal (1993, 1998), Montoya (1990, 1993, 1994, 1997), Azanza & Montoya (1995), Montoya & Alberdi (1995), Mom-