Journal of Physical Activity Research, 2018, Vol. 3, No. 2, 96-101
Available online at http://pubs.sciepub.com/jpar/3/2/6
©Science and Education Publishing
DOI:10.12691/jpar-3-2-6
Short-term Effects of Whey, Creatine, and L-carnitine
Supplementation on Muscle Hypertrophy Marker
Candidates in Young Males: A Randomized
Placebo-controlled Pilot Study
Bagus Sarmito
1
, Felicia Kartawidjajaputra
2,*
, Antonius Suwanto
1
1
Faculty of Biotechnology, Atma Jaya Catholic University of Indonesia, Jakarta, Indonesia
2
Nutrifood Research Center, PT Nutrifood Indonesia, Jakarta, Indonesia
*Corresponding author: felicia@nutrifood.co.id
Received August 13, 2018; Revised October 04, 2018; Accepted October 29, 2018
Abstract Previous research showed that resistance exercise could induce muscle development, noticeably from
the increased level of several markers from blood samples. However, no study had been performed to explore the
effect of combination of resistance exercise and proper nutrition supply on those markers. The aim of this study to
investigate the effect of whey protein, creatine, and L-carnitine; on potential molecular markers of muscle
hypertrophy (arg1 and mmp9) from blood samples. Twelve healthy male participants were randomly categorized
into supplement (SUPP) or placebo (PLAC) treatment, and performed resistance training three times in a one-week
period. Blood sampling was carried out before (day one) and 2 hours after the exercise (day one, day three and day
five). The level of mmp9 gene expression was increased along with the progress of the resistance training program.
Moreover, participants who received supplementation (SUPP) showed a higher level of mmp9 gene expression
compared to resistance training only (PLAC). A significant difference was observed between two treatments in the
first day, 2 hours after the resistance training session (p = .04); and between SUPP group on the fifth day, 2 hours
after the resistance training; compared to the first day, before the resistance training session (p = .02). The effect was
not observed on arg1 gene. A combination of resistance training with supplementation; was considered to enhance
the muscle hypertrophy process, compared to resistance training only. The results also suggested that mmp9 could
act as a blood-derived molecular marker of muscle hypertrophy.
Keywords: muscle hypertrophy, resistance training, whey protein, L-carnitine, mmp9, arg1
Cite This Article: Bagus Sarmito, Felicia Kartawidjajaputra, and Antonius Suwanto, “Short-term Effects of
Whey, Creatine, and L-carnitine Supplementation on Muscle Hypertrophy Marker Candidates in Young Males: A
Randomized Placebo-controlled Pilot Study.” Journal of Physical Activity Research, vol. 3, no. 2 (2018): 96-101.
doi: 10.12691/jpar-3-2-6.
1. Introduction
Resistance training is a kind of sport which can improve
general fitness status and induce muscular hypertrophy, by
performing movements against force in the opposite
direction. Studies showed positive effects of resistance
exercise to muscle cross-sectional area, cellular markers,
protein markers, and gene expression markers; yet, muscle
biopsy was still the method of preference for sampling
[1,2,3]. Muscle biopsy is known as an invasive method
and potential to cause a scar in the participant’s body.
Therefore, molecular markers from blood sample are
preferable because they give less invasive effect to the
participants.
Research showed that there were two genes which
upregulated in 2 hours after acute resistance exercise: arg1
and mmp9 [4]. Those genes were analyzed from blood
samples. Arginase-1 (Arg1) is an enzyme which
metabolizes arginine to ornithine, which furtherly
modified to proline, a constituent of collagen which
important in hypertrophy process [5,6,7]. Meanwhile,
matrix metalloproteinase 9 (MMP9, gelatinase B) is an
enzyme which is capable of degrading extracellular
matrices, particularly type IV and V collagen [8,9]. This
enzyme helps the movement of leukocytes toward the
damaged area of muscle tissue, improve the ability of
myoblast migration, and help the synthesis of new
myofiber [10,11]; thus, the hypertrophy process can occur.
The combination of resistance training and proper nutrition
supply will help the body-builders to achieve enhanced
muscular development. Administration of nutrients,
particularly whey protein, creatine, and L-carnitine would
further enhance the process of muscle hypertrophy.
Researches showed that whey protein and creatine could
improve myoblast proliferation and differentiation [12,13].
Meanwhile, L-carnitine was involved in muscle recovery