Pergamon
ft. Steroid Biochem. Molec. Biol. Vol. 52, No. 6, pp. 587-594, 1995
Copyright © 1995 Elsevier Science Ltd
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31p NMR Study of Phosphorus Containing
Metabolites in the Uterus of Hamster:
Changes During the Estrous Cycle and the
Effect of Hormonal Manipulation
S. Shivaji,* L. G. Devi, M. B. Ahmad and C. S. Sundaram
Centre for Cellular and Molecular Biology, Uppal Road, Hyderabad 500 007, India
Changes in the concelatrations of phosphorus containing metabolites were monitored by sip NMR
in the uteri of hamsters during the estrous cycle. Concentrations of phosphocreatine (PCr) and ATP
were significantly increased in estrus animals compared to diestrus animals. Concentrations of these
metabolites were also increased in immature female hamsters and ovariectomized (OVX) adult
hamsters treated with estradiol indicating that estradiol was responsible for this effect. However,
the steroid hormones; progesterone and testosterone did not increase the concentrations of the
phosphorus containing metabolites. Further, immature female hamsters also following treatment
with estradiol showed an initial decline in phosphomonoester (PME), PCr, ATP and inorganic
phosphate but by 24 h of treatment the concentrations returned to control levels. The NMR study
also revealed that the intracellular pH of the hamster uterus was around 7.4 all through the estrous
cycle.
J. Steroid Biochem. Molec. Biol., Vol. 52, No. 6, pp. 587-594, 1995
INTRODUCTION
In mammals, the uterus during the estrous cycle
exhibits cyclic morphological, biochemical and physio-
logical changes such as increase in tissue weight [1-3],
accumulation of fluid [4-,6] and increase in biosynthetic
activity [7]. Apart from an increase in the synthesis of
DNA, RNA and proteins under the influence of
estradiol, changes in the concentration of high energy
metabolites such as phosphocreatine (PCr) and
adenosine triphosphate (ATP) have also been observed
in the uterus of rat following estradiol administration
[8, 9]. But, all these earlier studies were performed
using perchloric acid extracts of frozen uteri thus
resulting in variations irt the reported results due to
partial degradation of the, labile compounds. However,
this difficulty could be overcome by the use of sip
NMR.
31p NMR has been extensively used for the analysis
of phosphate containing metabolites in excised tissues
and whole animals [10-12] so as to avoid extraction and
*Correspondence to S. Shivaji.
Received 7 Oct. 1994; accepted 26 Dec. 1994.
587
homogenization methods which are not suitable for the
analysis of labile compounds. Using this method, high-
energy phosphate metabolites in the uterus of adult rats
during pregnancy and parturition [13] and in the uteri
of immature rats following estradiol administration [14]
have been analyzed. However, as yet, very little is
known about the changes in the concentration of high-
energy phosphate metabolites during the functional
state of the uterus such as during the various phases of
the estrous cycle. Since steroid hormones play a
direct role in modulating the estrous cycle it would also
be important to evaluate the effect of steroid hormones
on the concentration of the high-energy phosphate
metabolites. Further, even less is known about the
intracellular pH of the uterus during the estrous cycle.
In an attempt to address the above questions the
present paper reports data related to:
(i) Changes in phosphate metabolites in adult
hamster uteri during diestrus, proestrus and
estrus phases of the estrous cycle.
(ii) Changes in phosphate metabolites in the uteri of
adult animals following ovariectomy and sub-
sequent administration of estradiol, progester-
one or testosterone.