© by PSP Volume 22 – No 12. 2013 Fresenius Environmental Bulletin
3567
ASSESSING INTRA-SPECIFIC VARIATION WITHIN
BACILLUS SPP. STRAINS USING DNA FINGERPRINTING
Ömür Baysal*
Muğla Sıtkı Koçman University, Faculty of Life Sciences, Department of Molecular Biology and Genetic, P.B. 48000 Kötekli Muğla, Turkey
ABSTRACT
Bacillus subtilis strains are known to be prominent
biologics playing different roles both in biological degra-
dation of organic compounds and bio-control of plant dis-
eases. In the present study, a schematic diagram was im-
proved using efficient ISSR markers, selected in previous
screening studies and suggested in view of their cost and
labor-effective roles in discrimination of Bacillus spp.
strains. To assess their efficiency and use in diversification
of Bacillus spp. strains, whole findings were compared
with 16S rDNA analysis. The results by selected Inter-
Simple Sequence Repeats (ISSR) markers showed conven-
ient, highly reproducible results to be a tool for revealing
genetic relationships in Bacillus spp. strains, with regard to
16S rDNA. Even ISSRs have no informative knowledge
based on gene sequence. Quick studies on structure and
dynamics of microbial communities in ecosystems are of
importance. Therefore, ISSR fingerprinting is suggested to
be an efficient tool in intraspecific discrimination of Bacillus
subtilis strains in further studies.
KEYWORDS: Molecular markers, intra-specific variation, 16S rDNA,
ISSR, Bacillus spp.
1. INTRODUCTION
Studies on the structure and dynamics of microbial
communities are relevant in microbial ecology to develop
bioremediation strategies. For identification of a microbial
agent, polymerase chain reaction (PCR) has been success-
fully used in different microbiological studies. Particularly,
the gene commonly amplified in such cases codes for 16S
rDNA, which is a highly conserved region both within a
species and among species of the same genus, and is essen-
tial for the survival of living organisms [1]. Therefore, 16S
rDNA analysis is also considered to be a standard method
for the identification of bacteria at family, genus and spe-
cies levels [2]. However, further discrimination of individ-
ual strains within a given species is needed in order to
understand the speciation and host preference occurrence
* Corresponding author
depending on pathogenic behavior. Consequently, a num-
ber of fingerprinting methods have been developed in
recent years to discriminate between strains belonging to
a given species, such as random amplification of polymor-
phic DNA [3] and amplified fragment length polymor-
phism (AFLP) analysis [4]. However, these methods are
not cost-effective. Inter-Simple Sequence Repeats (ISSR)
are dominant markers and detect polymorphisms in mi-
crosatellite and inter-microsatellite loci, and do not re-
quire prior information of DNA sequences [5]. In contrast
to other molecular markers, the target sequences for ISSR
primers help in revealing higher numbers of polymorphic
fragments than RAPD (Random Amplified Polymorphic
DNA) markers.
On the other hand, there is also no strict correlation
between function of Bacillus isolates and their genotypic
and phenotypic variations [6]. A relatively recent study
indicated that the function of a large number of genes in
B. subtilis remains unknown [7]. Most of the available evi-
dence concerning genotypic variation among different Ba-
cillus isolates has been acknowledged with assessment of
phenotypic variation and cell wall chemistry [8] but not
sufficiently informative on genus-based diversification.
In the present study, a schematic list using ISSR mark-
ers was developed for easy discrimination of Bacillus
strains to examine the species variability and to differenti-
ate closely related strains at the subspecies level, which
was also evaluated with the results obtained by 16S rDNA
analysis confirmations.
2. MATERIAL AND METHODS
2.1 Isolation, selection and identification of bacteria
In this work, 12 samples were isolated from green-
house soils collected during the Methyl Bromide Phase
Out Project of the Ministry of Agriculture in Turkey,
within 2003-2010. The bacterial suspensions were stored in
tubes containing King B medium at 4 ºC, and in flasks con-
taining TSB (tryptone soy broth) plus glycerol (20%, v/v) at
-20 ºC until use. Whole isolated bacteria were distinguished
from other B. subtilis colonies according to bacteriology
notes [9], including some tests. Afterwards, identification
of each strain was confirmed with 16S rDNA sequences