Physiology&Behavior. Vol. 54, pp. 753-758, 1993 0031-9384/93 $6.00 + .00
Printed in the USA. Copyright © 1993 Pergamon Press Ltd.
Role of Resocialization and of 5-HT1A Receptor
Activation on the Anxiogenic Effects Induced
by Isolation in the Elevated Plus-Maze Test
S. MAISONNETTE, S. MORATO AND M. L. BRANDAO l
Laboratory of Psychobiology, Faculdade de Filosofia, Ci~ncias, e letras de Ribeir~o Preto,
Campus-USP, 4040-901, Ribeir~o, Preto, S~o Paulo, Brazil
Received 14 January 1993
MAISONNETTE, S., S. MORATO AND M. L. BRANDAO. Role of resocialization and of 5-TH1A receptor activation on the
anxiogenic effects induced by isolation in the elevated plus-maze test. PHYSIOL BEHAV 54(4) 753-758, 1993.--Rats were
isolated for periods varying from 1 h to 2 weeks and the exploratory activity of these animals on the elevated plus-maze was
studied. Rats isolated from periods of 2 h on displayed a significant reduction in the number of entries and time spent in the
open arms of the plus-maze compared to socially housed controls. This effect was not correlated with the decrease in the total
number of entries also produced by isolation. Acute treatment with midazolam or resocialization for a 24-h period clearly reversed
these responses produced by prior 2-h isolation in the elevated plus-maze. It is suggested that exposure to a 2-h isolation period
could be a useful nonpharmacological means of generating anxiety in laboratory rodents. Chronic treatment, but not acute
treatment, with gepirone, a 5-HTtA agonist, inhibited the anxiogenic effects caused by a 2-week period of isolation. The reduction
in aversiveness promoted by resocialization may be due to a recovery in the 5-HT activity depressed by isolation in a much faster
way than observed with chronic gepirone administration.
Isolation Resocialization 5-HTIA receptors Gepirone Elevated plus-maze Aversion
ISOLATED rats demonstrate many behavioral disturbances such
as increased locomotor activity, neophobia, deficits in learning
involving or not spatial memory, and aggressive muricidal be-
havior (20,28,33). Neophobia and some of the other behavioral
differences observed in isolation-reared animals are thought to
be related to increased anxiogenic profile, and a raised level of
anxiety is observed when isolation-reared rats are placed in a
novel environment (23). Recently we have shown that the iso-
lation of rats for 2 weeks causes increased anxiety-related be-
havior in the elevated plus-maze test (21). In the present work
we were interested in examining what was the minimum period
of isolation capable of producing these anxiety effects in the
elevated plus-maze and also the minimal period ofresocialization
necessary to reverse them.
Previous research has suggested that brain serotonergic neu-
rotransmitter system participates in the modulation of anxiety
[for reviews see (13,14,16)], and that isolation is accompanied
by changes in the activity of this system (28,31). 5-HTIA agonists
like gepirone, as well as the benzodiazepines, all suppress neu-
ronal activity in the dorsal raphe nucleus following either sys-
temic or iontophoretic administration (3,1 1,30). Gepirone, which
binds with high affinity to the 5-HTIA receptor subtype (27), has
been reported to be clinically as effective as the standard ben-
zodiazepine, diazepam, as an anxiolytic (5). It has been shown
that upon acute administration the presynaptic effect ofgepirone,
which causes a reduction of 5-HT neuron activity, overrides its
postsynaptic action, an effect that results in a net decrease of
the serotonergic tone (19). However, it has been observed that
sustained administration of gepirone is necessary to attain a
clinically significant relief of anxiety (5). Based on these data,
we also carried out a comparative study of the effects ofgepirone,
acutely and chronically administered, and midazolam, a known
benzodiazepine agent, on the performance of socially isolated
rats in the elevated plus-maze.
METHOD
Animals
Male Wistar rats, weighing 220-250 g, were housed in groups
of six with free access to food and water. They were kept in
groups of six (except when stated otherwise) in cages of 60 × 60
× 30 cm for 72 h prior to the experiment, on a 12 h light:dark
cycle (lights on at 0700 h) in a room adjacent to the experimental
room. The temperature in both rooms was maintained between
25 and 27°C.
' Requests for reprints should be addressed to M. L. Brand~o.
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