Environmental Microbiology (2003) 5(8), 706–710
© 2003 Society for Applied Microbiology and Blackwell Publishing Ltd
Blackwell Science, LtdOxford, UKEMIEnvironmental Microbiology1462-2920Blackwell Publishing Ltd, 20035 8706710Short CommunicationPathogenic V. parahaemolyticus in the Seto Inland SeaM. J. Alam, S.-I. Miyoshi and S. Shinoda
Received 3 February, 2003; accepted 10 March, 2003. *For
correspondence. E-mail alammj30@hotmail.com; Tel. and Fax
(+81) 86 251 7967.
Brief report
Studies on pathogenic Vibrio parahaemolyticus during
a warm weather season in the Seto Inland Sea, Japan
M. Jahangir Alam,* Shin-Ichi Miyoshi and
Sumio Shinoda
Department of Environmental Hygiene, Faculty of
Pharmaceutical Sciences, Okayama University,
Tsushima, Okayama 700-8530, Japan.
Summary
Vibrio parahaemolyticus is a potentially pathogenic
bacterium, occurring naturally in estuarine and
marine environments throughout the world. The inci-
dence of this organism in an aquatic environment
depends upon many ecofactors. Sea water and
organic material were collected during the warm
weather season from a coast of the Seto Inland Sea,
Japan, and analysed to determine V. parahaemolyti-
cus densities and the occurrence of pathogenic
strains, defined as those possessing tdh and/or trh
genes by polymerase chain reaction (PCR), using iso-
lated DNA from enrichment culture of the samples.
About 99% of samples were positive for V. para-
haemolyticus with densities of 3 to > 1400 cells per
100 ml of water or 10 g of organic samples by the
most-probable-number (MPN)-PCR technique, but
only 76.6% were positive by the conventional MPN
culture technique, with densities ranging from 3 to
> 1400 cells per 100 ml of water or 10 g of organics.
Furthermore, the tdh and trh genes were positive in
41.5% and 8.5% of samples, respectively, by the MPN-
PCR technique. No tdh and trh gene-positive strains
were isolated by the conventional MPN culture proce-
dure. The difference in detection between the MPN
culture and the MPN-PCR techniques appeared to be
significant and may be attributed to different detec-
tion sensitivities and other factors.
The aquatic environment has been implicated as a prin-
cipal reservoir of many pathogenic vibrios. Vibrio para-
haemolyticus is a well-known human pathogen, causing
gastroenteritis through consumption of raw or under-
cooked seafood (Honda and Iida, 1993). This bacterium
has been found to account for about 20–30% of all food
poisoning cases in Japan. It is also reported to be an
important agent of travellers’ diarrhoea. Thermostable
direct haemolysin (TDH) and TDH-related haemolysin
(TRH), encoded by the tdh and trh genes, respectively,
are the best recognized pathogenic factors of V. para-
haemolyticus (Honda and Iida, 1993). Such pathogenic
factors have rarely been detected in environmental V.
parahaemolyticus isolates. The reason for this discrep-
ancy in virulence between clinical and environmental iso-
lates is not well understood.
Vibrio parahaemolyticus and many other vibrios become
abundant during warmer months in the sea waters of Japan
(Miyamoto et al., 1962; Chowdhury et al., 1990) and other
parts of the world but, in the cold weather months, they
are rarely isolated (Miyamoto et al., 1962; Kelly and Stroh,
1988; Chowdhury et al., 1990). The ecology and cultura-
bility of V. parahaemolyticus present in food and environ-
mental samples have not been fully understood.
Environmental conditions impose a variety of stresses on
bacteria, which in turn have developed different strategies
to survive, including adaptation and transmission. There-
fore, the incidence of a species in a water environment is
dependent upon many ecofactors. In environmental and
food samples, vibrios may be injured by various physico-
chemical factors and, therefore, may not grow in selective
and/or non-selective culture media. Furthermore, in
unfavourable environmental conditions, formation of via-
ble-but-non-culturable (VNC) cells has been reported for
more than 30 bacterial species (Xu et al., 1982; Colwell,
1993; Oliver, 1995; Oliver et al., 1995), but the role that
these cells might play in the environment and in patho-
genesis has so far not been resolved (Colwell, 1993;
Bogosian et al., 1998; Barer and Harwood, 1999). Cells
in the VNC state are unable to form colonies on plates but
are still alive and capable of metabolic activity (Colwell
and Huq, 1994). Moreover, VNC cells may remain poten-
tially pathogenic (Colwell and Huq, 1994), and can resume
active growth when environmental conditions are restored
(Colwell, 1993; Oliver, 1995; Whitesides and Oliver, 1997).
Detection of such cells requires alternative methods.