Eur. J. For. Path. 22 (1992) 284-290
© 1992 Verlag Paul Parey, Hamburg und Berlin
ISSN 0300-1237
Laboratorio di Botanica Forestale, Firenze, Italy
New type of damage to Quercus ilex:
pollen germination and hydration ability
By LoRENZA M. BELLANI and E. PAOLETTI
Abstract
Pollen grains collected from trees of Quercus ilex L. belonging to different classes of new type
damage were analyzed. Five parameters were taken into account: viability, germinability, tube
lengthening, grain volume of^ the dehydrated state and after rehydration. Damage to mother
plants clearly influences the above mentioned parameters, causing a reduction in pollen grain
viability, germinability, and in the ability of the grains to rehydrate.
1 Introduction
Several authors reported that the environment affects sexual reproduction, sex ratio being
shifted towards male or female individuals (HESLOP-HARRISON 1972; CHARNOV and BULL
1977; WiLLSON 1979). A tendency towards maleness in plant species is well known under
unfavorable conditions, e. g. drought (FREEMAN et al. 1976; LEV-YADUN and LIPHSCHITZ
1987) or elevational gradients (CRANT and MITTON 1979). CLAUSER et al. (1989) observed
a high production of male flowers in plants grown in urban and rural areas. This phenome-
non observed in plants of several species, such as Pinus sylvestris L., Cedrus atlantica Man-
etti, Cupressus sempervirens L. and mainly Quercus ilex L., was attributed to background
air pollution due to long range transport.
In damaged holm oak trees a change in leaf colour has been detected (CELLINI et al.
1988; CLAUSER et al. 1989). Leaves show a grayish colour of the foliage due to the twisting
of the leaf-stalk for which the abaxial green-gray surface instead of the dark green adaxial
one is shown. If the damage continues, the leaves of the previous and/or the current year
are shedded and secondary branch loss in the higher part of the crown is detected. At the
time of flowering damaged holm oak trees just keep newly emerged leaves and show a
great number of male flowers. When male flowers drop, the holm oak crown is extremely
scanty.
On the other hand, changes in reproductive processes caused by air pollution are not as
readily observed as are foliar injury symptoms (BENOIT et al. 1983).
In the present note we try to correlate viability, germinability and tube length in pollen
grains collected from Quercus ilex plants with different levels of new type of damage.
Simultaneously, the size of grains at the moment of release and their rehydration ability is
taken into account.
2 Materials and methods
2.1 Sample collection
Based on the Cerman tree damage classification (EEC 1987), plants are considered
healthy, with a loss of leaves inferior to 10 % (class 0), slightly damaged with a loss of
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