Vol.:(0123456789)
SN Applied Sciences (2019) 1:1538 | https://doi.org/10.1007/s42452-019-1560-7
Research Article
Monitoring skeletal muscle dynamics and modelling the nonlinear
response
M. Zakir Hossain
1,2,3
· Julian Grill
2
· Wolfgang Grill
1,2
Received: 2 July 2019 / Accepted: 22 October 2019
© Springer Nature Switzerland AG 2019
Abstract
A custom ultrasonic calliper was employed to monitor voluntary and externally excited muscle dynamics with synchro-
nous electromyography. The activation, hold, and relaxation phases of the gastrocnemius muscle were monitored for
maximum voluntary isometric contraction. Muscle belly shortening occurred during contraction and a post-contractile
overshoot (lengthening) and subsequent exponential recovery of muscle dimension to the baseline were observed.
Both the overshoot and recovery are attributed to the muscle as suggested by combined monitoring including elec-
tromyography and modelling with a lumped mechanical circuit containing idealized elements, such as a bidirectional
linear motor unit, a ratchet, dampers, and springs. The rapid contraction and relaxation phases require a high-order
filter or alternatively a kernel filter, attributed to the nervous system as suggested by external electric stimulation, which
resulted in faster rise and relaxation times. The respective response function is modelled with an electrical lumped
circuit. Together with empirically adjusted reaction times and corrections for droop in the hold phase, the monitored
response is represented in close approximation by the combined electrical and mechanical lumped circuits. The refined
combinatory model includes a ratchet as a novel nonlinear mechanical element. In combination with determined model
parameters, it provides a refined evaluation scheme capable to model monitored muscle dynamics in physical activity
in close approximation.
Keywords Ultrasonic monitoring of muscle dynamics · Modelling of muscle dynamics · Motor-spring-damper-ratchet
lumped circuit · Muscle response to external stimuli · Post-tetanus autonomous contraction
1 Introduction
Skeletal muscle dynamics have been an intriguing field of
study for centuries, having already been reported in 1667
[5, 18, 25]. Lumped circuit modelling of muscle dynamics
was introduced by Hill [8]. This model illustrated in Fig. 1a
and its adaptations have frequently been used to study
muscle properties, allowing advances in understanding of
fundamental dynamic properties. Recently more complex
models have been used in an attempt to model proper-
ties not suitably represented by the basic Hill model, like
the one proposed by Makssoud et al. [20] illustrated in
Fig. 1b. Hill’s historical three-element muscle model [7, 8]
has an active contractile element E, which is a linear motor
drive only capable of providing pull (shortening stress). It
has two springs S
1
and S
2
with S
1
in series with the con-
tractile element and S
2
parallel to both. These springs are
considered capable of transmitting pull only, resisting
elongation. Figure 1b represents the model proposed by
Makssoud et al. [20]. In addition to the classic contractile
and elastic elements, it contains two mass-spring-damper
arrangements. Both models connect to bones at each end
(grey boxes). Further historical perspectives of modelling
* Wolfgang Grill, wg@analogspeed.de |
1
Faculty of Physics and Earth Sciences, University of Leipzig, Leipzig, Germany.
2
ASI Analog Speed
Instruments GmbH, Burgweg 8, 61460 Koenigstein im Taunus, Germany.
3
Scientific Soccer School, Calgary, Canada.