Anim. Migr. 2020; 7: 19–26
https://doi.org/10.1515/ami-2020-0003
Received December 6, 2019; accepted February 24, 2020
Abstract: Cerulean Warblers (Setophaga cerulea) are
among the fastest declining Nearctic-Neotropical migrant
wood-warblers (Parulidae) in North America. Despite
ongoing conservation efforts, little is known about
their non-breeding distribution. In June 2016-2018, we
deployed geolocators (n = 30) on adult male Cerulean
Warblers in Indiana, USA, to track annual movements of
individuals. Recovered geolocators (n = 4) showed that
Cerulean Warblers occurred broadly throughout northern
South America. Autumn migration lasted 44-71 days (n =
4), whereas spring migration lasted 37-41 days (n = 3). The
average migration distance was 5268 km. During autumn
migration, Cerulean Warblers made 1-4 stopovers (i.e., ≥2
days; n = 4) and 1-2 stopovers during spring migration (n
= 3). When crossing the Gulf of Mexico during autumn
migration, two birds stopped over after crossing, but not
beforehand. Two others navigated through the Caribbean
rather than crossing the Gulf of Mexico. During spring
migration, one individual stopped after crossing, one
individual stopped before crossing, and one individual
stopped before and after crossing the Gulf of Mexico.
No birds migrated through the Caribbean Islands
during spring migration. These results represent novel
information describing annual movements of individual
Cerulean Warblers and will inform conservation efforts for
this declining species.
Keywords: full annual cycle, migratory connectivity,
Setophaga cerulea
1 Introduction
Identifying the geographic distribution and assortment of
migratory individuals during migration and nonbreeding
periods is critical for understanding population trends [1].
Populations may be limited by factors occurring outside
the breeding period including along population-specific
migratory routes [2] or at nonbreeding areas [3]. Until
recently, there has been a dearth of information on the
nonbreeding dispersion and migratory behaviors of most
species of New World wood-warblers (Parulidae). Most
parulids migrate considerable distances between Nearctic
breeding grounds and Neotropical nonbreeding grounds
[4], and many of these species are declining [5]. The
recent miniaturization of tracking technology (light-level
geolocators) has made tracking small migratory species (<
10 g) possible [6-8].
Cerulean Warblers (Setophaga cerulea) are small
(~9 g), Nearctic-Neotropical migratory songbirds that
breed in mature deciduous forests of the eastern United
States and adjacent southern Canada. Individuals
overwinter in northern South America along the eastern
slopes of the northern Andes at elevations of ~500-
2000 m [9]. Cerulean Warblers are among the fastest
declining Nearctic-Neotropical migrant wood-warbler
species in North America (~3% per year) [9] with declines
purportedly linked to extensive habitat loss on both
breeding and nonbreeding grounds [9, 10]. However,
because Cerulean Warbler research, conservation, and
management are focused on the breeding grounds, there is
currently little information on the dispersion of Cerulean
Warbler populations outside of the breeding period
(i.e., stopovers during autumn and spring migrations,
and the nonbreeding period), which may be hindering
conservation efforts if limiting factors are experienced
away from the breeding grounds [3]. Determining where
populations of Cerulean Warblers overwinter, the routes
they use to migrate between breeding and nonbreeding
*Corresponding author: Kamal Islam, Ball State University, Muncie,
IN United States, E-mail: kislam@bsu.edu
Clayton D Delancey, Garrett J MacDonald, Alexander R Sharp, Bran-
don M Connare, Ball State University, Muncie, IN United States
Gunnar R Kramer, University of Toledo, Toledo, OH United States
Clayton D Delancey, Kamal Islam*, Gunnar R Kramer, Garrett J MacDonald, Alexander R Sharp,
Brandon M Connare
Geolocators reveal migration routes, stopover
sites, and nonbreeding dispersion in a population
of Cerulean Warblers
Communication
Open Access. © 2020 Kamal Islam et al., published by De Gruyter. This work is licensed under the Creative Commons Attribution 4.0 Public
License.