© 2019 Academia Sinica, Taiwan
Open Access
Age, Growth, and Sex Ratios of the Giant
Mottled eel, Anguilla marmorata, in Freshwater
Habitats Near Its Northern Geographic Limit: A
Comparison to Tropical Regions
Ryoshiro Wakiya
1,
*, Hikaru Itakura
2
, and Kenzo Kaifu
3
1
Research and Development Initiative, Chuo University, 1-13-27 Kasuga, Bunkyo-ku, Tokyo 112-8551, Japan.
*Correspondence: Tel: +818056758725. E-mail: ryoshiro.wakiya@gmail.com
2
Graduate School of Sciences, Kobe University, 1-1 Rokkoudaichou, Nadaku, Kobe, Hyogo 657-8501 Japan, E-mail: itakurahikaru@gmail.com
3
Faculty of Law, Chuo University, 742-1 Higashinakano, Hachioji-shi, Tokyo 192-0393 Japan, E-mail: kaifu@tamacc.chuo-u.ac.jp
Received 19 March 2019 / Accepted 18 September 2019 / Published 12 November 2019
Communicated by Benny K.K. Chan
To understand the latitudinal diference in the basic ecology of the giant mottled eel Anguilla marmorata,
we measured its age, growth, and sex ratios in freshwater areas of Amami-Oshima Island, Japan
(28.223°N–28.332°N and 129.329°E–129.439°E), near the northern geographic limit of its range and
compared these biological characteristics with those observed in the tropical regions of Indonesia. A total
of 109 A. marmorata were captured from three rivers on Amami-Oshima Island. The total length (TL) and
age of sampled individuals varied across an order of magnitude (TL range: 119–1320 mm, mean: 385.5 ±
172.6; age range: 3–30 years, mean: 12.8 ± 4.9 years). Neither TL nor age difered among rivers. Male A.
marmorata accounted for 88.5% of sexually diferentiated individuals, which it contrary to previous results
for males living in Indonesian watersheds where males were only found in small rivers and areas close to
estuaries. Moreover, we found no males larger than 700 mm in TL and few males that were older than 20
years. The growth rates of male A. marmorata were signifcantly lower than those of females, and growth
rates of sexually undiferentiated fsh were signifcantly less than those of males. The mean growth rate
of all individuals was 25.9 ± 6.6 mm/y, which is considerably lower than what has been found at lower
latitudes, suggesting that growth diferences occur along a latitudinal cline.
Key words: Age, Anguilla marmorata, Freshwater, Growth rate, Sex ratio.
Citation: Wakiya R, Itakura H, Kaifu K. 2019. Age, growth, and sex ratios of the giant mottled eel, Anguilla marmorata, in freshwater habitats near
its northern geographic limit: a comparison to tropical regions. Zool Stud 58:34. doi:10.6620/ZS.2019.58-34.
BACKGROUND
Species in the genus Anguilla consist of
catadromous fish that contain leaf-like leptocephalus
larvae that migrate from open oceans to freshwater and
estuarine habitats (Han et al. 2016; Chen et al. 2018;
Higuchi et al. 2018). After metamorphosing into yellow
eels, the fish spend most of their life in continental
waters until they begin their downstream migration
toward spawning grounds in open ocean. The anguillid
eel consists of 19 species and subspecies distributed
globally, with the exception of polar regions and the
west coast of North and South America (Ege 1939;
Aoyama 2003; Watanabe et al. 2004). The giant mottled
eel A. marmorata is the most broadly distributed species
in the genus, with a range from the south-east African
coast to Asia and Polynesia (Ege 1939; Marquet and
Galzin 1991; Marquet 1996). It is also one of the largest
species in the genus, growing up to 2 m and 21 kg (Castle
1984). Despite this prominent ecology, to date, little
basic biological information is available on the giant
mottled eel, likely because compared with the Japanese
Zoological Studies 58: 34 (2019)
doi:10.6620/ZS.2019.58-34
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