1
Volume 7 / Issue 1 / 9 / http://dx.doi.org/10.21931/RB/2022.07.01.9
Optimization of cephalosporin C acylase immobilization using crosslinked
enzyme aggregates technique
Julkipli Julkipli
1
, Khaswar Syamsu
1
and Ahmad Wibisana
2
*
Abstract: Cephalosporin C acylase (CCA) is an essential enzyme for the one-step conversion of cephalosporin C into
7-aminocephalosporanic acid (7-ACA), an intermediate compound used to synthesize various semi-synthetic cephalosporin
antibiotics. The industrial process prefers to use enzymes in immobilized form rather than soluble. A crosslinked enzyme
aggregate (CLEAs) is a potential matrix-less enzyme immobilization technique to produce stable immobilized enzymes with
high activity and low production costs. This study aimed to optimize the CCA immobilization using the CLEAs technique with
Chitosan as a co-aggregate. The CCA lysate was obtained from harvesting CCA fermentation broth using a mutant strain of
Escherichia coli through cell separation and lysis steps. Partially purified CCA by ammonium sulfate addition was conducted to
obtain an active fraction of 20-60% saturation, followed by co-aggregation with Chitosan to form physical CCA aggregates. The
aggregates were then immobilized by a crosslinking technique using glutaraldehyde to form CLEAs-CCA. Optimization of the
immobilization process was carried out by Response Surface Methodology in three steps, (i) screening of the influencing factors,
(ii) determining the level of the significant factors, and (iii) optimizing the immobilization condition. The CLEAs-CCA activity was
used as a response parameter. Under optimum conditions, CLEAs-CCA activity obtained was 85.91 Ug
-1
.
Key words: 7-aminocephalosporanic acid, cephalosporin C acylase, Chitosan, crosslinked enzyme aggregates, response surface.
methodology.
ARTICLE / INVESTIGACIÓN
Introduction
Cephalosporins are antibiotics with a broad antibacterial
spectrum and almost cover 50% of β-lactam antibiotics hu-
mans use
1,2
. Most of them are derived from 7-aminocephalos-
poranic acid (7-ACA), a precursor compound generated from
hydrolysis of cephalosporin C (CPC), a natural antibacterial
obtained from Acremonium chrysogenum fermentation
3
. The-
refore, it is essential to produce 7-ACA efficiently.
The industrial demands for efficient and environmentally
friendly 7-ACA production have shifted from chemical to en-
zymatic processes. Initially, a two-step bioconversion using
D-amino acid oxidase and glutaryl-7-aminocephalosporanic
acid acylase was developed
4
. Recently, one-step bioconversion
using cephalosporin C acylase (CCA, EC 3.5.1.11) has paid the
attention of researchers because it is more efficient
5
.
The industrial application of enzymes is preferred in the
immobilized rather than soluble form due to the more efficient
bioconversion process
6
. Immobilization of enzymes on a matrix
is a popular method. However, a dilution of enzyme up to 99%
occurred, resulting in decreased activity
7
. In addition, the ma-
trix used causes an increase in the enzyme cost and generates
a waste matrix generated from the inactive biocatalyst
8
. To
overcome these problems, enzyme immobilization by cross-
linked enzyme aggregates (CLEAs) technique, a matrix-less
immobilization approach, is a potential method to be applied
7
.
It showed a simple, fast, and efficient method to immobilize
various enzymes
8-10
. To the best of our knowledge, there is no
publication on the application of CLEA for CCA immobilization.
The CLEAs formation involves two critical steps, i.e., en-
zyme precipitation to form aggregates and crosslinking of the
aggregates
7
. Ammonium sulfate and glutaraldehyde are often
used as precipitation and crosslinker agents, respectively
9,11,12
.
However, crosslinking the enzyme with low superficial lysine
residues with glutaraldehyde may cause weak enzyme bin-
ding, making it quickly released from its CLEAs
13
. The addition
of co-aggregates, such as bovine serum albumin, poly-ethyle-
neimine, and poly-lysine, could form larger aggregates size
and higher stability
9,12,13
. Chitosan is a relatively inexpensive
biopolymer (polyaminosaccharides, copolymers of N-ace-
tyl-D-glucosamine and N-amino-D-glucosamine) with high
availability rich in reactive amino groups, potentials to mediate
the formation crosslinks between the aggregates
14,15
.
In this research, optimization of the CCA immobilization by
CLEAs technique using Chitosan as a co-aggregate was stu-
died. Optimization was conducted with Response Surface Me-
thodology (RSM), as it is an effective and widely used method
for optimization in bioprocesses, including enzyme immobili-
zation
16-18
. CLEAs-CCA (Ug-1) activity was used as a response.
The experiments were carried out in three steps, namely, (i)
screening of the influencing factors using 2
n-2
fractional fac-
torial design (FFD), (ii) determining the level of significant fac-
1
Biotechnology Study Program, Graduate School, IPB University, IPB Dramaga Campus, Bogor, Indonesia.
2
The National Research and Innovation Agency (BRIN), Tangerang Selatan, Banten, Indonesia and Chemical Engineering Study Program, Pamulang University,
Witana Campus, Tangerang Selatan, Banten, Indonesia.
Corresponding author: ahmad.wibisana@brin.go.id
DOI. 10.21931/RB/2022.07.01.9
Citation: Julkipli J, Syamsu K and Wibisana A. Optimization of cephalosporin C acylase immobilization using crosslinked enzyme aggregates
technique. Revis Bionatura 2022;7(1). 9. http://dx.doi.org/10.21931/RB/2022.07.01.9
Received: 13 June2021 / Accepted: 10 August 2021 / Published: 15 February 2022
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