Abstract—This paper presents a framework for controlling
the development of a vascular system in an in vitro angiogenesis
process. Based on on-line measurement of cell growth and a
stochastic model of cell population, a closed-loop control system
is developed for regulating the process of cell migration and
tissue formation. Angiogenesis develops in a wet environment
and it is difficult to control each and every cell individually and
specifically. Instead, chemical and mechanical stimuli can be
applied pervasively to the whole process as global control
inputs, which can allow for control of collective behaviors of the
cell population. This paper formulates a systems level
description of the angiogenesis process and proposes a control
scheme that chooses global control inputs to drive collective cell
patterns, such as branch density per unit length of sprout,
toward a desired goal. In response to control inputs, the k-step
ahead prediction of collective cell pattern is evaluated, and the
input that is most likely to bring the predicted cell pattern to the
desired one is selected for the current control. Simulation
demonstrates that the global branch density of a simplified
angiogenesis model can be controlled using this technique. To
our knowledge, this paper is the first to formulate feedback
control for regulating an in vitro angiogenesis process.
I. INTRODUCTION
NGIOGENESIS is the process of growing or extending a
vascular network into a tissue matrix from a preexisting
vascular system. Understanding how to control angiogenesis
is extremely important in many research areas, ranging from
cancer treatment and wound healing to morphogenesis, stem
cells, and tissue engineering. During angiogenesis,
endothelial cells (EC’s) sprout from existing vasculature and
extend into the surrounding tissue. The sprouts extend and
branch as a function of local chemical and mechanical
conditions. The angiogenesis process has not been
completely described via deterministic models and cellular
behaviors have been described as stochastic processes
Manuscript received April 23, 2008. This material is based on work
supported in part by the National Science Foundation under grant number
NSF EFRI-0735997.
L. B. Wood is with the department of mechanical engineering at the
Massachusetts Institute of Technology, Cambridge, MA 02139 USA
(phone: 617-258-8013; e-mail: woodl@mit.edu).
A. Das is with the department of biological engineering at the
Massachusetts Institute of Technology, Cambridge, MA 02139 USA (e-
mail: anusuya@mit.edu).
R. D. Kamm is with the department of biological engineering and the
department of mechanical engineering at the Massachusetts Institute of
Technology, Cambridge, MA 02139 USA (e-mail: rdkamm@mit.edu).
H. H. Asada is with the department of mechanical engineering at the
Massachusetts Institute of Technology, Cambridge, MA 02139 USA (e-
mail: asada@mit.edu).
modulated by conditions local to the cell [1],[2]. Regulating
and manipulating the angiogenesis process by actively
modulating chemical and mechanical conditions is a truly
challenging research issue, which will have a significant
impact upon broad biological engineering and medical fields.
Angiogenesis and other biological processes are
fundamentally different from traditional engineering systems
where control technology has been successfully applied.
First, the system consists of a vast number of cells that have
local controllers to perform a specific class of functions.
Collective behavior of the cells exhibits meaningful
functionality, such as constructing a vascular network.
Second, cells are living in a “wet” environment, where
signals propagate through diffusion. Stimuli to the process
affect broad regions of cells.
In designing a control architecture, it is important to
understand that it is not fundamentally possible to control the
behavior of each and every cell in the population. The
available stimulus is much too limited in specificity. Further,
it is not necessary, or even desirable, to control each and
every cell. In generating a vascular network, the system as a
whole should satisfy certain collective requirements, such a
vascular density in the matrix, rate of branching, rate of
growth, or others. The behavior of each individual cell does
not matter, but correct development of the cellular
population as a whole is important in growing a useful
vascular system.
An additional important feature of the angiogenesis
process is that system stimuli do not effect deterministic
changes in system behavior. Instead, application of a
stimulus modulates cell transition probabilities. The authors
have previously developed stochastic broadcast feedback
control for control of clouds of independent cells and vast
degree of freedom actuators [3],[4]. The authors foresee that
insights gained from controlling the meaningful collective
behavior of the angiogenesis process will yield insight into
better design of vast degree of freedom systems like robotic
swarms through engineered randomness.
The objective of this paper is to develop a control
framework for regulating collective measures of vascular
growth. We will pose a systems level description of the
angiogenesis process with global outputs that we wish to
control. Inspiration for the systems level description must be
obtained from mathematical models of the angiogenesis
process. In addition, implementing control will require
detailed angiogenesis models. It may also be necessary to
utilize chemical diffusion models to describe the dynamics
A Stochastic Control Framework for Regulating Collective
Behaviors of an Angiogenesis Cell Population
Levi B. Wood, Anusuya Das, Roger D. Kamm and H. Harry Asada, Member, IEEE
A
Proceedings of the 2nd Biennial IEEE/RAS-EMBS International
Conference on Biomedical Robotics and Biomechatronics
Scottsdale, AZ, USA, October 19-22, 2008
978-1-4244-2883-0/08/$25.00 ©2008 IEEE 390