Journal of Radioanalytical and Nuclear Chemistry, Vol. 255, No. 1 (2003) 149–153
0236–5731/2003/USD 20.00 Akadémiai Kiadó, Budapest
© 2003 Akadémiai Kiadó, Budapest Kluwer Academic Publishers, Dordrecht
Water movement in a plant sample by neutron beam analysis
as well as positron emission tracer imaging system
T. M. Nakanishi,
1
* Y. Okuni,
1
J. Furukawa,
1
K. Tanoi,
1
H. Yokota,
1
N. Ikeue,
2
M. Matsubayashi,
2
H. Uchida,
3
A. Tsiji
3
1
Graduate School of Agricultural and Life Sciences, The University of Tokyo, Tokyo, 113-8657, Japan
2
Japan Atomic Energy Research Institute, Tokai Establishment, 319-1195, Japan
3
Hamamatsu Photonics, Co., Hamamatsu, 434-8601, Japan
(Received April 18, 2002)
We present water imaging of a plant sample both by neutron beam and positron emission tracer imaging system (PETIS). The former method
provided static water profile in a plant sample as well as that in the vicinity of a root imbedded in soil. Not only X-ray film but also CT method
using a cooled CCD camera is presented. Through non-destructive water image in an X-ray film, root development as well as 2-dimensional water
movement toward the root was analyzed. Spatial water image was constructed from 180 CT projection images, taken at an interval of one degree
while rotating the sample, through a CCD camera. In the case of a soybean root, there was a water gradient toward a root in soil and gave minimum
value at about 1 mm far from the surface of a root. The water absorbing part in a root was gradually shifted downward with the root development.
We also present real time water movement by PETIS, where water was labeled with a positron emitting nuclide,
15
O. The transportation of
15
O-
water within a plant was relatively slow and water uptake was observed only at the lowest internode, between a root and the first leaf, during
20-minute measurement.
Introduction
Though water plays an important role in plant
physiology, the behavior of water has not been known
well because of tools lacking for the research. Although,
the imaging by X-ray beam is an effective technique to
visualize comparatively heavy elements in living
samples. Neutron beam has an advantage to detect light
elements, such as hydrogen, lithium and boron. When
the plants are irradiated with neutrons beam, hydrogen
image is produced, for the attenuation efficiency of
hydrogen is about 100–1000 times higher compared to
those of the other common elements. Since more than
80% of living organism is consisted of water, an image
by neutron beam can be regarded as water image of a
living plant. Recently, NAKANISHI et al.,
1-8
has
developed neutron beam analysis to get water image of a
living plant. This method provides nondestructive water
specific image in a plant with high resolution. We also
performed a neutron beam imaging through a cooled
CCD camera to construct CT images (computer
tomography; cross-sectional image), and spatial water
distribution was analyzed according to the method
developed. Though the neutron method showed the
static water distribution in a living plant, it was difficult
to trace the real-time water movement in a plant tissue.
Therefore, we tried a method, namely the positron
emitting tracer imaging system (PETIS), where the
radioactive nuclide emits two gamma-rays, with
identical energy, to 180 degree. Setting the sample
between a pair of gamma-ray detector, the position and
amount of the labeled water absorbed can be measured.
* E-mail: atomoko@mail.ecc.u-tokyo.ac.jp
First of all, we tried to use
18
F-labeled water
(T
1/2
= 110 min), produced by a cyclotron, to measure a
short term water uptake and transport in a plant,
9-11
which method was developed by KUME et al.
12
However, there was always a problem whether tracing
an activity from
18
F really indicates the water movement
itself. Though only a small amount of
18
F was produced
from water target, without any carrier of F
-
ions, the
movement of
18
F
-
ions could be different from that of
water itself. To solve this problem, the application of
15
O-labeled water was necessary. But because of the
extremely short half-life of
15
O (2 min), it was difficult
to produce enough amount of tracer and to perform the
tracer work. Using the special system developed by one
of the authors, we could produce sufficient amount of
15
O-water to measure the real time movement of water
itself in a soybean plant. In this paper we present the
water image obtained by neutron beam as well as PETIS
using
15
O-water.
Experimental
Neutron imaging
Soybean seedlings (Glycine max c.v. Tsurunoko)
were grown in an aluminum container
(7 cm 15 cm 2 mm) containing Toyoura’s standard
sand (silt type, 197–203 m, in pore size) with 15%
(w/w) of water. The sample was grown in a phytotron at
27 °C and was periodically taken out to get neutron
image. Neutron irradiation was performed at a research
reactor, JRR-3M at the Japan Atomic Energy Research