Research Article
Diversity of Leptospira spp. in Rats and Environment from
Urban Areas of Sarawak, Malaysia
Chai Fung Pui,
1
Lesley Maurice Bilung,
1
Kasing Apun,
1
and Lela Su’ut
2
1
Department of Molecular Biology, Faculty of Resource Science and Technology, Universiti Malaysia Sarawak,
94300 Kota Samarahan, Sarawak, Malaysia
2
Faculty of Medicine and Health Sciences, Universiti Malaysia Sarawak, 94300 Kota Samarahan, Sarawak, Malaysia
Correspondence should be addressed to Lesley Maurice Bilung; mblesley@unimas.my
Received 11 November 2016; Revised 9 January 2017; Accepted 6 February 2017; Published 28 February 2017
Academic Editor: Jean-Paul J. Gonzalez
Copyright © 2017 Chai Fung Pui et al. Tis is an open access article distributed under the Creative Commons Attribution License,
which permits unrestricted use, distribution, and reproduction in any medium, provided the original work is properly cited.
Various prevalence studies on Leptospira in animals and humans, as well as environmental samples, had been conducted worldwide,
including Malaysia. However, limited studies have been documented on the presence of pathogenic, intermediate, and saprophytic
Leptospira in selected animals and environments. Tis study was therefore conducted to detect Leptospira spp. in rats, soil, and
water from urban areas of Sarawak using the polymerase chain reaction (PCR) method. A total of 107 rats, 292 soil samples, and
324 water samples were collected from April 2014 to February 2015. Pathogenic Leptospira was present in 5.6% (6/107) of rats, 11.6%
(34/292) of soil samples, and 1.9% (6/324) of water samples. Intermediate Leptospira was present in 2.7% (8/292) of soil samples and
1.9% (6/324) of water samples. Saprophytic Leptospira was present in 10.3% (11/107) of rats, 1.4% (4/292) of soil samples, and 0.3%
(1/324) of water samples. From this study, 76 Leptospira spp. were isolated. Based on DNA sequencing, the dominant Leptospira
spp. circulating in urban areas of Sarawak are pathogenic Leptospira noguchii, intermediate Leptospira wolfi serovar Khorat, and
saprophytic Leptospira meyeri, respectively. Overall, this study provided important surveillance data on the prevalence of Leptospira
spp. from rats and the environment, with dominant local serovars in urban areas of Sarawak.
1. Introduction
Leptospirosis is a zoonotic disease with worldwide distribu-
tion and caused by pathogenic Leptospira, which results in
signifcant public health problem worldwide [1, 2]. Te genus
Leptospira consists of 20 species, with more than 300 serovars,
grouped into 20 serogroups [3]. Based on the pathogenic-
ity, they can be divided into three major clades, namely,
pathogenic, saprophytic (nonpathogenic), and intermediate
(unclear pathogenicity) [2]. Te pathogenicity status of inter-
mediate Leptospira remains a debate matter. For instance,
hamsters inoculated with intermediate L. inadai and L. licera-
siae do not cause any clinical manifestation of leptospirosis
although both recovered from patients [4].
Humans usually get infected through direct contact with
the infected animal urine via mucous membrane and exposed
skin or indirect contact by exposure to the contaminated
soil, water, and food [5–7]. Normally, maintenance hosts are
asymptomatic while accidental hosts like humans may sufer
a wide range of clinical manifestation such as renal failure,
hepatic failure, severe pulmonary haemorrhage, and even
death [8]. Te overall case mortality rate in humans ranges
from 1 to 5% and the elderly are claimed to have higher infec-
tion risk of leptospirosis [9]. Incidence rates are ofen under-
estimated because of the relative inaccessibility, lack of rapid
diagnostics, and insufcient awareness of leptospirosis [10].
Similar prevalence studies on Leptospira spp. had been
conducted in Malaysia in recent years. Leptospira had been
isolated from urban rats of Kuala Lumpur [11] as well as soil
and water from urban sites in Peninsular Malaysia [12]. Te
presence of Leptospira spp. was also reported in National Ser-
vice Training Centres of Peninsular Malaysia [13]. Moreover,
Tayaparan et al. [14] reported the presence of Leptospira spp.
in wildlife around tourism areas in Sarawak. Our previous
study highlighted the presence of Leptospira spp. in national
parks [15]. Te presence of Leptospira spp. in this area posed
the risk of transmission and infection to humans. Since this
study on the detection of Leptospira spp. in rats, soil and
Hindawi
Journal of Tropical Medicine
Volume 2017, Article ID 3760674, 8 pages
https://doi.org/10.1155/2017/3760674