Citation: Tsai, S.-C.;Huang, Y.-W.;
Wu, C.-C.; Wang, J.-J.; Chen, Y.-T.;
Singhania, R.R.; Chen, C.-W.;
Dong, C.-D.; Hsieh, S.-L.
Anti-Obesity Effect of Nostoc
commune Ethanol Extract In Vitro and
In Vivo. Nutrients 2022, 14, 968.
https://doi.org/10.3390/
nu14050968
Academic Editor: Lindsay Brown
Received: 13 January 2022
Accepted: 22 February 2022
Published: 24 February 2022
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nutrients
Article
Anti-Obesity Effect of Nostoc commune Ethanol Extract In Vitro
and In Vivo
Sheng-Chieh Tsai
1,2
, Yu-Wen Huang
1
, Chih-Chung Wu
3
, Jyh-Jye Wang
4
, Ya-Ting Chen
1
,
Reeta Rani Singhania
5
, Chiu-Wen Chen
5
, Cheng-Di Dong
5
and Shu-Ling Hsieh
1,
*
1
Department of Seafood Science, National Kaohsiung University of Science and Technology,
Kaohsiung 81157, Taiwan; b220682837@gmail.com (S.-C.T.); may2377234@gmail.com (Y.-W.H.);
melodyyu.chen@gmail.com (Y.-T.C.)
2
Department of Medical Research, E-Da Hospital, Kaohsiung 82445, Taiwan
3
Department of Food and Nutrition, Providence University, Taichung 43301, Taiwan; wuccmail@gmail.com
4
Department of Nutrition and Health Science, Fooyin University, Kaohsiung 83102, Taiwan; ft054@fy.edu.tw
5
Department of Marine Environmental Engineering, National Kaohsiung University of Science and
Technology, Kaohsiung 81157, Taiwan; reetasinghania@nkust.edu.tw (R.R.S.);
cwchen@nkust.edu.tw (C.-W.C.); cddong@nkust.edu.tw (C.-D.D.)
* Correspondence: slhsieh@nkust.edu.tw; Tel.: +886-7-361-7141 (ext. 23626)
Abstract: Nostoc commune is an edible terrestrial blue-green alga. It has shown many beneficial effects
on human health. This study aimed to investigate the phytochemical assay of N. commune ethanol
extract (NEE) and its anti-obesity effects. The effect of a high-calorie diet on lipid accumulation in
3T3-L1 preadipocytes is investigated, and a Wistar rat model is used to demonstrate the anti-obesity
effect of NEE and its mechanism. The results showed that the NEE has phytochemical compounds,
such as total polyphenol, total flavonoids, and total terpenoids. NEE was also shown to suppress cell
proliferation and lipid accumulation (26.9%) in 3T3-L1 preadipocytes. Furthermore, NEE reduced the
body weight (13.5%), fat tissue weight (13.3%), and the serum FFA (19.4%), TG (14.2%), TC (11.8%),
and LDL-C (16.4%) of rats. In histopathology, NEE was shown to diminish the size of adipocytes
and hepatic lipid droplets. The NEE downregulated the mRNA expression of adipogenesis (PPAR-γ,
SREBP-1c) and lipid lysis-related genes (ATGL, HSL) in epididymal adipose tissue. The NEE also
upregulated the mRNA expression of β-oxidation related genes (AMPK, CPT-1, PPAR-α) in the
liver. Overall, this study suggests NEE has the potential to be developed as a functional food for
anti-obesity.
Keywords: Nostoc commune; phytochemical; anti-oxidations ability; 3T3-L1 pre-adipocytes; obesity
1. Introduction
Obesity is defined as an excess of fatty tissue in the body, which increases the risk of a
variety of chronic illnesses and body function disorders, such as diabetes mellitus type 2,
hypertension, cardiovascular disease, and obese sarcopenia. [1,2]. Adipocyte size and quan-
tity increase, resulting in a low-grade pro-inflammatory milieu in adipose tissue, which
leads to fat gain [3]. It is important to study the role of fat in obesity for the prevention as
well as treatment of obesity. Lipogenesis and lipolysis are important processes for the con-
trol of adipocyte energy balance. Peroxisome proliferator-activated receptor γ (PPARγ) is
one of the transcription factors that is important in the transformation of preadipocytes into
mature adipocytes; sterol regulatory element-binding protein (SREBP) expression helps
triglyceride synthesis in adipocytes, which leads to adipocyte lipid accumulation [4,5].
In addition, a decrease in adipocyte size is an effective therapy for obesity. Adenosine
monophosphate-activated protein kinase (AMPK) is a key factor of energy homeostasis,
because it is a fuel-sensing enzyme that is activated by an increased ratio of AMP/ATP
and is sensitive to nutrition conditions [6,7]. Hormone-sensitive lipase (HSL) and adipose
Nutrients 2022, 14, 968. https://doi.org/10.3390/nu14050968 https://www.mdpi.com/journal/nutrients