Vol.:(0123456789) 1 3
https://doi.org/10.1007/s10096-022-04413-8
REVIEW
Resolving taxonomic confusion: establishing the genus Phytobacter
on the list of clinically relevant Enterobacteriaceae
Theo H. M. Smits
1
· Lavinia N. V. S. Arend
2
· Sofa Cardew
3
· Erika Tång‑Hallbäck
4
· Marcelo T. Mira
5
·
Edward R. B. Moore
6,3
· Jorge L. M. Sampaio
7
· Fabio Rezzonico
1
· Marcelo Pillonetto
2,5
Received: 2 November 2021 / Accepted: 25 January 2022
© The Author(s) 2022
Abstract
Although many clinically signifcant strains belonging to the family Enterobacteriaceae fall into a restricted number of genera
and species, there is still a substantial number of isolates that elude this classifcation and for which proper identifcation
remains challenging. With the current improvements in the feld of genomics, it is not only possible to generate high-quality
data to accurately identify individual nosocomial isolates at the species level and understand their pathogenic potential but
also to analyse retrospectively the genome sequence databases to identify past recurrences of a specifc organism, particularly
those originally published under an incorrect or outdated taxonomy. We propose a general use of this approach to classify
further clinically relevant taxa, i.e., Phytobacter spp., that have so far gone unrecognised due to unsatisfactory identifcation
procedures in clinical diagnostics. Here, we present a genomics and literature-based approach to establish the importance
of the genus Phytobacter as a clinically relevant member of the Enterobacteriaceae family.
Keywords Phytobacter diazotrophicus · Phytobacter ursingii · Phytobacter palmae · Genomics · Identifcation · Taxonomy
Introduction
The taxonomy of Enterobacteriaceae, since its establish-
ment and valid publication in the Approved Lists of Bacte-
rial Names in 1980 [1], has been prone to constant updates,
revisions and corrections [2, 3]. Over the years, many novel
genus and species names have been validly published. Since
about 1985, the use of improved molecular tools such as PCR,
16S rRNA gene and multi-locus sequence analysis (MLSA)
has led to a more stable taxonomy [4]. It has been only in
the last 10 years that genome-based taxonomic studies and
the use of average nucleotide identities (ANI) [5] and digital
DNA-DNA hybridization (dDDH) [6] have enabled reliable
delineation and detailed analysis of the diferent taxa, at the
species-level, and giving rise in 2016 to the revision of the
Enterobacteriaceae, with the creation of several sister fami-
lies, like the Erwiniaceae or Pectobacteriaceae [2]. A more
detailed analysis has shown six distinct phylogenomic-based
* Theo H. M. Smits
theo.smits@zhaw.ch
* Marcelo Pillonetto
m.pilonetto@pucpr.br
1
Environmental Genomics and Systems Biology Research
Group, Institute of Natural Resource Sciences (IUNR),
Zurich University of Applied Sciences ZHAW, Wädenswil,
Switzerland
2
Central Public Health Laboratory – State of Paraná
– LACEN/PR, Molecular Bacteriology Division,
São José Dos Pinhais, PR, Brazil
3
Culture Collection University of Gothenburg (CCUG),
Department of Clinical Microbiology, Sahlgrenska
University Hospital, Region Västra Götaland, Gothenburg,
Sweden
4
Department of Clinical Microbiology, Sahlgrenska
University Hospital, Region Västra Götaland, Gothenburg,
Sweden
5
Core for Advanced Molecular Investigation, Graduate
Program in Health Sciences, School of Medicine, Pontifícia
Universidade Católica Do Paraná, Curitiba, PR, Brazil
6
Department of Infectious Disease, Institute for Biomedicine,
Sahlgrenska Academy, University of Gothenburg,
Gothenburg, Sweden
7
Faculdade de Ciências Farmacêuticas - University of São
Paulo and Fleury Medicina Diagnóstica, São Paulo, SP,
Brazil
/ Published online: 15 February 2022
European Journal of Clinical Microbiology & Infectious Diseases (2022) 41:547–558