pubs.acs.org/JAFC Published on Web 06/17/2010 © 2010 American Chemical Society
J. Agric. Food Chem. 2010, 58, 8085–8092 8085
DOI:10.1021/jf101325m
New Mathematical Model for Interpreting pH-Stat Digestion
Profiles: Impact of Lipid Droplet Characteristics on
in Vitro Digestibility
YAN LI AND DAVID JULIAN MCCLEMENTS*
Biopolymers and Colloids Research Laboratory, Department of Food Science,
University of Massachusetts, Amherst, Massachusetts 01003
The pH-stat method is commonly used to characterize the in vitro digestibility of lipids under
simulated small intestine conditions. This method measures the fraction of free fatty acids (FFA)
released from triacylglycerols over time. A new mathematical model has been developed to
characterize the FFA versus time profiles generated by the pH-stat method, which can be used
to quantify the influence of physicochemical parameters on the rate (k) and extent (φ
max
) of lipid
digestion. In this model, k is the amount of FFA produced per unit time per unit surface area,
whereas φ
max
is the maximum fraction of digestible FFAs released. This model is used to quantify
the influence of lipid droplet characteristics (size, concentration, composition, and emulsifier type) on
the digestion of emulsified lipids. The rate (k) of lipid digestion increased with decreasing lipid
content (from 2.5 to 0.5 wt %), increasing droplet diameter (from d = 200-15000 nm), and
decreasing fatty acid molecular weight (MCT versus corn oil). The extent (φ
max
) of lipid digestion
was also considerably less for corn oil than for MCT. The rate and extent of lipid digestion did not
depend strongly on initial emulsifier type: β-lactoglobulin, Tween 20, lecithin, or lyso-lecithin. These
results are interpreted in terms of differences in the concentrations of reactants, products, catalysts
and cofactors at the lipid droplet surfaces during digestion, for example, triacylglycerols, emulsifiers,
FFA, lipase, and bile salts. This model provides a useful means of quantifying the influence of
specific parameters on lipid digestion using the pH-stat method.
KEYWORDS: Emulsions; lipid digestion; lipase; calcium; calcium binding; pH-stat
INTRODUCTION
There is growing interest in understanding and controlling the
digestibility of emulsified lipids as they pass through the human
gastrointestinal (GI) tract ( 1 -5 ). This knowledge is being used by
the pharmaceutical industry to rationally design emulsion-based
delivery systems to carry biologically active agents to specific
locations within the GI tract and release them at a controlled
rate ( 6 , 7 ). Analogous systems are also being developed by the
food industry to encapsulate, protect, and deliver bioactive food
components (nutraceuticals) ( 4 , 5 , 8 -11 ). These delivery systems
have been proposed as a means of increasing the bioavailability of
highly lipophilic nutraceuticals or as a means of delivering
functional food ingredients to specific locations within the GI
tract, such as the colon for chemopreventative agents. Emulsion-
based delivery systems with controlled stability and digestibility
within the GI tract have also been proposed to be an effective
method of controlling appetite and therefore combating
obesity ( 12 -16 ). In this case, studies have found that emulsions
which remain stable to gravitational separation in the stomach
and/or which have a delayed digestion in the small intestine can
stimulate the release of gut hormones that induce satiety and
reduce food intake. Consequently, there is great interest in the
development of analytical tools that can be used to establish the
major physicochemical and structural factors that affect lipid
digestion and absorption under conditions that simulate the
human GI tract. These tools range from physicochemical mea-
surements using in vitro digestion tests, to cell culture models, to
animal feeding studies, and ultimately to human trials ( 4 , 17 , 18 ).
The pH-stat method is an analytical tool that is finding
increasing use within pharmaceutical and food research for the
in vitro characterization of lipid digestion under simulated small
intestinal conditions ( 19 -22 ). The pH-stat method is based on
measurements of the amount of free fatty acids (FFAs) released
from lipids, usually triacylglcyerols (TAGs), after lipase addition
at pH values close to neutral. The lipid-containing sample to be
analyzed is placed within a temperature-controlled reaction
chamber that contains simulated small intestinal fluid (SSIF).
The SSIF typically includes appropriate concentrations of the
major digestive components known to influence lipid digestion,
such as lipase, colipase, bile salts, phospholipids, and mineral
ions. The lipase in the SSIF catalyzes lipid digestion leading to the
generation of two FFAs and one monoacylglycerol (MAG) per
TAG molecule. The concentration of alkali (NaOH) that must
then be titrated into the digestion cell to neutralize any FFA
produced by lipid digestion, and thereby maintain the pH at the
*Corresponding author [e-mail mcclements@foodsci.umass.edu;
phone (413) 545-1019; fax (413) 545-1262].