American-Eurasian J. Agric. & Environ. Sci., 11 (3): 371-384, 2011
ISSN 1818-6769
© IDOSI Publications, 2011
Corresponding Author: El-Refaey F.A. El-Dengawy, Department of Pomology, Faculty of Agriculture, Damietta Branch,
Mansoura University, Damietta, Egypt.
371
Improving Growth and Salinity Tolerance of
Carob Seedlings (Ceratonia siliqua L.) by Azospirillum Inoculation
El-Refaey F.A. El-Dengawy, Ahmed A. Hussein and Saad A. Alamri
1,2 2 2
Department of Pomology, Faculty of Agriculture,
1
Damietta Branch, Mansoura University, Damietta, Egypt
Department of Biology, Faculty of Science, King Khalid University, Abha, Saudi Arabia
2
Abstract: Carob has been neglected with respect to both cultural practices and research and development. The
current study was carried out for two successive seasons to find a practical method for extension of carob
cultivation under saline water irrigation and to evaluate the efficacy of the Azospirillum inoculation in the
development of salinity tolerance of the carob seedlings under salinity stress (4.69EC, 9.38EC and 14.07EC of
seawater). The results showed that the seedling growth characteristics particularly seedling dry weight, the new
leaf area and root characters showed significant progressive reductions with increasing salinity levels. But
these reductions were significantly ameliorated by Azospirillum inoculation. Regarding the leaf mineral and
proline content, the saline irrigated seedlings either inoculated or non-inoculated with Azospirillum contained
significantly higher Na , Cl and proline concentrations and significantly lower K and N concentrations in leaf
+ - +
than that of the normal irrigated control. However, applying Azospirillum inoculum in combination with saline
water significantly decreased Na+ and Cl- and increased K+, N and proline concentrations in the seedling
leaves under all salinity levels. Also, K/Na ratio was significantly increased by Azospirillum inoculum under
the most tested levels of salinity. Applying Azospirillum inoculum gave significantly higher total phenols
concentration than uninoculated seedlings. The SCC percentage was significantly higher in saline water
irrigated seedlings at 14.07EC with or without Azospirillum inoculum than the other tested treatments.
Chlorophyll concentration in leaves of the treated seedling was significantly decreased with rising levels of
salinity. Applying Azospirillum inoculum increased chlorophyll concentration in seedling leaves under all
salinity levels compared to the uninoculated ones. Moreover, applying Azospirillum inoculum in combination
with saline water significantly increased survival ratio of the seedlings and decreased injured leaves (%)
compared to the uninoculated ones under the same level of salinity. It can be conclude that application of
Azospirillum inoculum can initiate systemic resistance of carob seedling to salt stress and improve growth.
Key words:Carob seedlings Salinity stress Azospirillum Seedling characteristics Physiological
parameters
INTRODUCTION Azospirillum was discovered as a growth promoter of
The carob (Ceratonia siliqua L.) is found in a range conducted on cereal, the perception of Azospirillum as a
of the wild habitat of many Mediterranean countries plant-growth-promoting bacterium (PGPB) for cereals is
including Cyprus, Egypt, Italy, Lebanon, Libya, southern still widespread. This ignores the multitude of studies in
Jordan, Syria, Tunisia and Turkey, representing different which Azospirillum affected and promoted the growth of
genetic resources [1]. The origin of carob is not clear as it numerous other species, including trees, cacti, vegetables,
has undergone extensive cultivation since ancient times fruit, flowers, medicinal plants and spices [5]. This review
[2]. Carob is also widely planted as an ornamental and proposed that Azospirillum be considered a non-specific
shade tree in some countries as USA and Australia [3]. PGPB. Successful applications and research in these areas
Besides, all parts of the carob tree, e.g. pods, seeds and are progressing slowly and most studies are conducted
woods have great economic aspects [4]. under in vitro conditions. Research in these areas is
cereals. Because much Azospirillum research is still