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Phyto-toxic Efect of Heavy Metal (CdCl
2
) on Seed Germinaton, Seedling Growth and Antoxidant
Defence Metabolism in Wheat (Tritcum aestvum L.) Variety HUW-234
Anaytullah Siddique
*
and Anand Prakash Dubey
Dept. of Plant Physiology, Insttute of Agricultural Sciences, RGSC, Banaras Hindu University, Varanasi, U.P. (221 005), India
The experiments were conducted to assess the phytotoxic efect of heavy metal (CdCl
2
). The responses of treatment efect comprises the
variables germinaton %, length (cm), fresh weight and dry weight (mg) of shoot and root, root/shoot rato, seed vigour index (SVI), reducing
sugar, non reducing sugar, total sugar (mg g
-1
), chlorophyll content (mg g
-1
), cadmium uptake (ppm g
-1
), proline content (µg g
-1
), relatve
water content (RWC%) and superoxide dismutase (SOD, unit×10
2
g
-1
f. weight min
-1
). CdCl
2
treated sets showed reducton in percent of seed
germinaton, length (cm), fresh weight and dry weight (mg) of shoot and root, root/shoot rato, seed vigour index (SVI) with increasing the
concentratons of CdCl
2
in comparison to control set. A reducton in breakdown of reducing sugar to non reducing sugar and total sugar is
correlated with decline of amylase actvity along with increasing the concentratons of CdCl
2
in comparison to control one. The chlorophyll
content was found in decreasing trend with increasing the concentratons of CdCl
2
in comparison to control. However, the uptake of
cadmium was increased with increasing the concentratons up to 100 ppm CdCl
2
. The proline content and superoxide Dismutase (SOD)
was found to be high in higher concentratons of CdCl
2
treated set and while the enhancement was started since inital concentraton CdCl
2
(i.e., 25 ppm). However, the least amount was recorded in control set. On the contrary, RWC% was found to be reduced with increasing
the concentratons of CdCl
2
up to 100 ppm even though the maximum RWC% was measured in control set.
1. Introducton
Wheat (Tritcum aestvum L.) belongs to family Gramineae and
is one of the leading cereals of many countries in respect to
growth, producton and consumpton of the world (Farooq
et al., 2011a). India is the world’s second largest producer
of wheat (Goncalves et al., 2011). Thus, it is important
to study the toxicity of heavy metals on this crop. Metal
contaminaton is always increasing by the actvites of human
which interferes with the environment and makes adverse
conditon for living organism. Some plant species have capacity
to grow in the metal contaminated soil and accumulate
high amount of heavy metals (hyper-accumulaton) as an
eco-physiological adaptaton in Metalliferous soil. Among
the contaminatons heavy metals Pb, Cd, Hg, As, Ni, and
Se are emerging problems for all over worlds. (Ahmad and
Ashraf, 2011; Ahmad et al., 2011a). Cadmium (Cd) is one of
the most common toxic metals present in the environment
that induces various toxic efects in plants even at low doses.
Contaminaton of soil in cultvated felds with toxic heavy
metals such as cadmium, copper, nickel and zinc has emerged
as a new challenge to agriculture (Singh et al., 2007). Inital
development of the plant, comprises various processes of seed
germinaton (Dominguez and Cejudo, 2014) that culminates
the metabolic transformatons that trigger the development
of the embryonic axis, consequently, the emergence of radical
(Oliveira et al., 2013). On the other hand, heavy metals are
abiotc factors that may interfere in the development of seed
germinaton. The presence of cadmium permanently in the
soil contaminates water and impedes the absorpton of plant
nutrients causing morphological, physiological, biochemical
and structural changes in plants (Augusto et al., 2014). The
immobilizaton of cadmium outside of the plant or inside of
the root may be one of the frst barriers of the plant against
toxicity to the heavy metal since cadmium levels are highest
in roots rather than in leaves, with low transport level from
the root to the aerial segment of the plant (Chaves and Souza,
2014). Although, several studies about Cd toxicity to the wheat
have been reported but stll it is needed to explore Cd toxicity
in wheat crop (Shaf et al., 2010; Rizwan et al., 2016).
2. Materials and Methods
A lab experiment was conducted in the Department of Plant
Heavy metal, CdCl
2
, wheat, SOD, SVI, RWC%, proline Keywords:
Abstract
Article History
Manuscript No. AR1684
Received in 9
th
September, 2016
Received in revised form 19
th
February, 2017
Accepted in fnal form 19
th
March, 2017
Anaytullah Siddique
e-mail: anaytullahsiddique@gmail.com
Corresponding Author
261
International Journal of Bio-resource and Stress Management 2017, 8(2):261-267
Doi: HTTPS://DOI.ORG/10.23910/IJBSM/2017.8.2.1684
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