Volume 2 • Issue 1 • 1000108
Enz Eng
ISSN: EEG, an open access journal
Research Article Open Access
Zhao et al., Enz Eng 2013, 2:1
DOI: 10.4172/2329-6674.1000108
Research Article Open Access
Keywords: Fish waste; Pepsinogen; Extraction; Purifcation;
Fractionation; Enzyme activity; Protein concentration; Recovery yield
Introduction
Atlantic Canada has 15% (40,000 km) of the Canada’s coastline
and 42% of the Canada’s fresh water (320,000 km
2
). About 80% of total
fsh landings come from the Atlantic fshery, while the Pacifc fshery
accounts for up to 16% [1]. In 2010, Atlantic Canada’s total sea fsheries
landings were 788,599 t/y, with the total value of 1.3 billion dollars. A
large portion of the fsh (80%) landed in Atlantic Canada is processed
[2].
Tere are three major common steps in fsh processing: (a)
removing the viscera, (b) removing the head, tail, fns and skin, and
(c) removing the frame and producing fllets. A large fraction (30-80%)
of fsh (fesh, heads, bones, fns, skin, tails and viscera) is generated as
waste during fsh processing [3,4]. Fish wastes are usually disposed of in
landflls or poured directly into the sea, which results in high disposal
cost and causes environmental problems. About 56% of the fsh
landing in Canada is converted into waste; of which 13% is disposed
in the ocean [5]. Conventional disposal of fsh wastes underscores the
need for a more reasonable utilization approach of fsh wastes, as well
as efective recovery of valuable ingredients from these wastes.
Fish wastes can be utilized as animal feed ingredients, as well as
organic fertilizers [6,7]. Te recovery of valuable biomolecules, such
as collagen [8-11], ω-3fatty acids [12], trypsin [13,14], chymotrypsin
[15-17], and elastase [18], have also been reported. Among the valuable
products that can be recovered from fsh, pepsin is one of the abundant
and useful biomolecules that can be efectively recovered from fsh
viscera.
Pepsin is synthesized and secreted in the gastric membrane in an
inactive state called Pepsinogen (PG) (molecular weight of 40 kDa).
Pepsin is an important acidic protease, widely applied in the hydrolysis
of proteins in the food and manufacturing industries. It can be used
in collagen extraction [19-21], gelatin extraction [22], cheese making
[23], and regulating digestibility [24]. Extraction of pepsins from
fsh viscera not only signifcantly reduces the capital costs of enzyme
production, but also partially improves the economics of the fsh
processing industry, while minimizing the environmental impact of
waste disposal [25,26].
Pepsin, as well as its zymogen, pepsinogen (PG), has been widely
purifed from several fsh species, including arctic fsh capelin (Mallotus
villosus) [27], rainbow trout (Salmo gairdneri) [28], Atlantic cod
(Gadus morhua) [29], bolti fsh (Tilapia nilotica) [7], Antarctic rock
cod (Trematomus bernacchii) [30], sea bream (Sparuslatus houttuyn)
[31], African coelacanth (Latimeria chalumnae) [32], Mandarin fsh
*Corresponding author: Abdel E Ghaly, Department of Process Engineering and
Applied Science, Dalhousie University, Halifax, Nova Scotia, Canada, Tel: 902-
494-6014; E-mail: abdel.ghaly@dal.ca
Received March 01, 2013; Accepted March 02, 2013; Published March 10, 2013
Citation: Zhao L, Budge SM, Ghaly AE, Brooks MS, Dave D (2013) Partition of
Pepsinogen from the Stomach of Red Perch (Sebastes marinus) by Aqueous Two
Phase Systems: Effects of PEG Molecular Weight and Concentration. Enz Eng 2:
108. doi: 10.4172/2329-6674.1000108
Copyright: © 2013 Zhao L, et al. This is an open-access article distributed under
the terms of the Creative Commons Attribution License, which permits unrestricted
use, distribution, and reproduction in any medium, provided the original author and
source are credited.
Abstract
Fish processing waste can be used to produce commercially valuable by-products, such as pepsinogen, which
has application in food, manufacturing industries, collagen extraction, gelatin extraction, and in regulating digestibility.
An important acidic protease, pepsin, is synthesized and secreted in the gastric membrane in an inactive state called
pepsinogen (PG). In the present study, the purifcation of pepsinogen from the stomach of red perch, using aqueous
two phase systems (ATPS) formed by polyethylene glycol (PEG) and salt at 4
°
C, was optimized.The effects of PEG
molecular weight (PEG 1000, 1500, 3000 and 4000) and concentration (16, 18, 20, 22 and 24%) on the partitioning
of PG were studied, and parameters including total volume (TV), volume ratio (V
R
), total enzyme activity (A
E
), protein
content (C
p
), specifc enzyme activity (SA), partition coeffcient (K
p
), purifcation fold (PF), and recovery yield (RY)
were evaluated. PEG molecular weight and PEG concentration also had signifcant effects on each parameter. TV
and V
R
decreased with increased salt concentration, since salt formed hydrogen bonds with water molecules and
formed a more compact and ordered water structure. PG partitioned predominantly in the PEG-rich top phase due
to its negative charge. A
E
, C
P
, SA, PF and RY increased with increased salt concentration and then decreased,
while K
P
had an opposite pattern. The PEG 3000 (20%), PEG 1000 (24%), PEG 4000 (16%) and PEG 1000 (18%)
concentrations gave the highest TV, V
R
, C
P
and K
P
, respectively. PEG 1500 with 18% concentration gave the highest
A
E,
SA, PF and RY (86.2%)
.
As PEG 1500 at 18% concentration gave the highest RY (86.2%). It was selected as
the optimum PEG molecular weight and PEG concentration. (NH
4
)
2
SO
4
at 15%, which gave the highest RY (71.7%),
was selected as the optimum salt type and salt concentration. 15% (NH
4
)
2
SO
4
18% PEG 1500 was the optimal ATPS
combination, and presented a better partition. The values of SA and PF and RY obtained with ATPS method were
much higher (2 fold in case of SA and PF, and 1.2 fold in case of RY), than those obtained with the Ammonium
Sulphate Fractionation (ASF) method.
Partition of Pepsinogen from the Stomach of Red Perch (Sebastes
marinus) by Aqueous Two Phase Systems: Effects of PEG Molecular
Weight and Concentration
Lisha Zhao, Suzanne M Budge, Abdel E Ghaly*, Marianne S Brooks and Deepika Dave
Department of Process Engineering and Applied Science, Faculty of Engineering, Dalhousie University, Halifax, Nova Scotia, Canada
E
n
z
y
m
e
E
n
g
i
n
e
e
r
i
n
g
ISSN: 2329-6674
Enzyme Engineering