Aquatic Invasions (2010) Volume 5, Supplement 1: S1-S4
This is an Open Access article; doi: 10.3391/ai.2010.5.S1.001
© 2010 The Author(s). Journal compilation © 2010 REABIC
Open Access
S1
Aquatic Invasions Records
First record of the Japanese shore crab Hemigrapsus sanguineus (de Haan, 1835)
(Brachyura: Grapsoidea: Varunidae) from the Black Sea
Drago ş Micu
1*
, Victor Ni ţă
1
and Valentina Todorova
2
1
National Institute for Marine Research and Development “Grigore Antipa”, 900581 Constanţa, Romania
2
Institute of Oceanology of the Bulgarian Academy of Sciences “Fridtjof Nansen”, Varna, Bulgaria
E-mail: dragos.micu@gmail.com (DM), victor_nicolae@yahoo.com (VN), vtodorova@io-bas.bg (VT)
* Corresponding author
Received: 10 November 2009 / Accepted: 4 December 2009 / Published online: 11 January 2010
Abstract
The Japanese shore crab Hemigrapsus sanguineus is recorded for the first time from a Black Sea locality: Tomis Marina in
Constanţ a, Romania. The suggested vector of introduction is as adults in the hull fouling of yachts. The species has not
established and salinity requirements for larval development make it unlikely that it will ever establish in the Black Sea.
Key words: Hemigrapsus sanguineus, Black Sea, Romania, not established
Introduction
The Japanese shore crab Hemigrapsus
sanguineus (de Haan, 1835) is native to the
North-Western Pacific, where it is distributed
from Peter the Great Bay, Russia to Hong Kong,
including the coasts of Japan, Korea, China and
Taiwan (McDermott 1998a). It lives in high
energy habitats, usually in crevices between
boulders of rocky shores, but also on sand
provided some rocks are present (Fukui 1988;
Benson 2005).
The species already has a history of
transoceanic invasions. In September 1988
H. sanguineus was first discovered in New
Jersey, on the Atlantic coast of the United States
and by 1990 a breeding population was already
established (Williams and McDermott 1990;
McDermott 1991). At present it ranges from
Maine to North Carolina (Delaney et al. 2008).
In 1999 it was recorded from the Atlantic
coasts of Europe, in Le Havre, France and the
Oosterschelde estuary in The Netherlands
(Breton et al. 2002). It has established there and
now is present in Germany, The Netherlands,
Belgium and France (Obert et al. 2007;
d’Udekem d’Acoz and Faasse 2002; Faasse
2004; d’Udekem d’Acoz 2006; Kerckhof et al.
2007; Dauvin 2009; Dauvin et al. 2009).
In 2003 a single adult male Hemigrapsus
sanguineus was recorded from the northern
Adriatic Sea (Schubart 2003) and this remains
the only known Mediterranean record to date.
Material and methods
In August 2008 we studied the rocky midlittoral
and sublittoral crustacean fauna around
Constanţa, Romania by snorkeling and SCUBA
diving. Samples were fixed in 10% buffered
seawater-formalin for 24 h and then transferred
to 70% ethanol for storage.
During this investigation one adult male
Hemigrapsus sanguineus was collected inside
Tomis Marina (N 44°10'43'', E 28°39'37'') on 27
August 2008, from under a rock just above the
waterline (Figures 1-2). The specimen has been
deposited at the National Museum of Natural
History “Naturalis”, Leiden, under accession
number RMNH D 53141.
Intense efforts to catch additional specimens
were unsuccessful in spite of the large extent of
the searches. Tomis Marina was subsequently
surveyed bimonthly by one of the authors, from
August 2008 to October 2009. The search was
extended to the adjacent commercial harbour
Constanţa-Sud Agigea and other harbours along
the Romanian coast (Midia Harbour, Eforie Ma-
rina, Mangalia Harbour) which were examined
repeatedly over this same period. No other
specimens of H. sanguineus were found.