Research Article
Evaluating the Biodeterioration Enzymatic Activities of
Fungal Contamination Isolated from Some Ancient Yemeni
Mummies Preserved in the National Museum
Khalid Mohammed Naji,
1
Qais Yusuf M. Abdullah,
2
Aida Qaseem M. AL-Zaqri,
3
and Saeed M. Alghalibi
2
1
Department of Chemistry, Faculty of Science, Sana’a University, Sana’a, Yemen
2
Department of Biology, Faculty of Science, Sana’a University, Sana’a, Yemen
3
Department of Archaeology, Faculty of Art, Sana’a University, Sana’a, Yemen
Correspondence should be addressed to Khalid Mohammed Naji; khalid.m.naji@gmail.com
Received 29 July 2014; Revised 25 September 2014; Accepted 16 October 2014; Published 13 November 2014
Academic Editor: Saad Tayyab
Copyright © 2014 Khalid Mohammed Naji et al. Tis is an open access article distributed under the Creative Commons Attribution
License, which permits unrestricted use, distribution, and reproduction in any medium, provided the original work is properly cited.
Sophisticated mummifcation using chemical preservation was prevalent in ancient Yemeni civilization as noted in the 4th century
B.C. mummies of the National Museum of Yemen, Sana’a, used in this study. Five of these mummies were used to evaluate hydrolytic
enzymes produced as a result of fungal contamination. Forty-seven fungal species were isolated, thereby refecting a high degree of
contamination which may have resulted from the poor ventilation and preservation system. Aspergillus was the most common genus
isolated (48.9%). Fifeen isolates exhibited ability to produce cellulase (EC; 3.2.1.4), Aspergillus candidus being the highest cellulose-
producer. Pectin lyase (PL, EC; 4.2.2.2) and pectin methyl esterase (PME, EC; 3.1.1.11) were produced by Trichoderma hamatum,
whereas chitinase (EC; 3.2.1.14) was produced by Aspergillus niger. Protease activity was noted by only Cladosporium herbarum. Te
higher activities of these fungal hydrolytic enzymes represent the major threats of biodeterioration including deteriorating linen
bandages as well as the mummy bodies. Terefore, it is recommended to improve the preservation system of the mummies at the
National Museum to minimize the contamination up to the lowest level and protect the mummies from biodeterioration.
1. Introduction
Human mummies are extremely susceptible to damage from
environmental factors [1]. Tere are a number of abiotic and
biotic factors, such as pollution, light, humidity, tempera-
ture, microorganisms, and insects, that have deteriorating
efects on museum materials [2]. Te most serious damage
on mummies occurs by microorganisms such as bacteria,
actinomycetes, and fungi [1, 3, 4].
In an indoor environment, fungi are the most hazardous
microorganisms responsible for the biocontamination of
museum collections. Unlike bacteria whose growth in muse-
ums is limited by their need for water, fungi are able to grow
at relatively low levels of temperature and humidity such as
those usually maintained in the museums [5, 6]. Even in quite
harsh conditions including dryness and low temperature,
fungal spores are able to survive for a long time [7]. Fungi are
usually present in display and storage conditions of museums
[8, 9].
Fungal degradation is one of the highest risk factors for
deterioration of ancient mummies [10, 11]. Te deterioration
by fungi seems to be a predominant feature in the museums
of many countries all over the world [8, 12]. Fungi excrete
enzymes that digest organic matter, thereby altering and
weakening those materials. In addition, many molds contain
colored substances that can cause stains and spots on textiles
and decrease the strength of the fabric. Many fungi can
be dangerous to people and in some cases pose a major
health hazard and cause zoonotic superfcial infections as a
consequence of invading keratinized tissues of skin, hair, and
nails [13, 14].
Fungal biodeterioration results in a change in the quality
or value of a material and makes it less functional in uti-
lization terms. A battery of extracellular hydrolytic enzymes
Hindawi Publishing Corporation
Biochemistry Research International
Volume 2014, Article ID 481508, 9 pages
http://dx.doi.org/10.1155/2014/481508