Vol.:(0123456789) 1 3
International Journal of Peptide Research and Therapeutics
https://doi.org/10.1007/s10989-018-9709-6
Optimization of Human Serum Albumin Periplasmic Localization
in Escherichia coli Using In Silico Evaluation of Diferent Signal Peptides
Mohammad Hassan Jahandar
1
· Ali Forouharmehr
2
Accepted: 13 April 2018
© Springer Science+Business Media, LLC, part of Springer Nature 2018
Abstract
Signal peptides (SPs) are essential tools to keep folding and function of recombinant proteins which are folded by disulfde
bonds. Human serum albumin (HSA) is a protein with medical application which is folded by 17 disulfde bonds. Therefore,
the production of this protein as recombinant protein needs appropriate SP. This study was performed to identify the best SP
to express HSA in Escherichia coli. At frst, 42 signal sequences (SSs) were collected from the database, then SP probability
and SP regions were predicted by SignalP (version 4). Physico-chemical features of SPs were also evaluated by Portparam,
whereas signal solubility was assessed by SOLpro. Finally secretion sorting of SPs were investigated by PRED-TAT server.
The results of this study revealed that, among 42 signal sequences, FlgI, DsbG, OmpC and MepA (respectively) are theoreti-
cally the best SPs to express HSA in E. coli.
Keywords Human serum albumin · Signal peptide · E. coli · Bioinformatics
Introduction
Human serum albumin (HSA) is one of the most abun-
dant proteins in serum and plasma which is released by
liver cells. Molecular mass of this protein is 67.25 kDa
with 591 amino acids and it has three domains which are
folded by 17 disulfde bonds. Half-life of HAS is around 20
days and has an important role in the adjustment of blood
osmatic pressure. These features of HSA are considered, to
apply this protein as therapeutic molecule for the treatment
of patients with burn injuries (Colmenarejo 2003; Leich
et al. 2007; Peters Jr 1995; Rosenoer et al. 2014). For this
purpose, nowadays, HSA is being produced through blood
plasma extraction, moreover, this method is too expensive
and also increases the risk of pathogenic pollutions. There-
fore, the production of HSA as recombinant protein can be
an appropriate alternative for the currently used methods.
HSA is a simple protein which does not need post transla-
tion modifcation; hence, this protein (as a recombinant
protein) can be easily expressed in prokaryote expression
system like Escherichia coli (Peters Jr 1995). Although
prokaryote expression system is simple and afordable
in expressing recombinant proteins, this system requires
some considerations such as maintaining disulfde bonds
(Ali et al. 2017). As matter of fact, in order to maintain
disulfide bonds and recombinant protein folding, they
must be directed to periplasm compartment of Bacteria,
because this compartment, in contrast to cytoplasm, can
provide reductive condition to save disulfde bonds and
consequently to fold the recombinant proteins. To direct
a nascent recombinant protein to periplasm compartment,
a signal peptide (SP) is required (Ali et al. 2017). In fact,
SP is an amino acid sequence in the N-terminal of nascent
proteins with three important regions including: n, h and c
regions (respectively) (Ali et al. 2017; Manica et al. 2017).
n Region has positive charge that helps it to interact with
the membrane of periplasm compartment; h region usually
has hydrophobic features and is important in translocation
of nascent proteins across membrane between cytoplasm
and periplasm; whereas c region helps SP to be cleaved
from recombinant protein (Ali et al. 2017; Zamani et al.
2015). Although SPs play an important role in expressing
recombinant protein, unfortunately there is no universal
* Mohammad Hassan Jahandar
Jahandar@iaubam.ac.ir
Ali Forouharmehr
Forouharmehr@gmail.com
1
Department of Agriculture, Bam Branch, Islamic Azad
University, Bam, Iran
2
Department of Animal Science, Faculty of Agriculture,
Lorestan University, Khorramabad, Iran