Variation of Prokaryotic and Fungal Soil
Communities across a Vegetative Transect
Sarah E Shawver
1
, Jeff A Brady
2
and Donald G McGahan
1,2
*
1
Department of Wildlife, Sustainability and Ecosystem Sciences, Tarleton State University, USA
2
Texas A&M AgriLife Research, USA
Submission: March 09,2018; Published: April 11, 2018
*
Corresponding author: Donald G McGahan, Department of Wildlife, Sustainability, and Ecosystem Sciences, Tarleton State University, Box T-0050,
Stephenville, TX 76402, USA, Email:
Agri Res & Tech: Open Access J 15(3): ARTOAJ.MS.ID.555953 (2018) 0069
Research Article
Volume 15 Issue 3 - April 2018
DOI: 10.19080/ARTOAJ.2018.15.555953
Agri Res & Tech: Open Access J
Copyright © All rights are reserved by Donald G MCGahan
Introduction
Soil microbial communities are affected by a number of
biotic and abiotic factors including pH, nitrogen (N), carbon
(C), and vegetation. Soil pH is one of the primary influences
on microbial communities. Soil chemical properties, especially
pH, rather than spatial effect, land use, climate, or soil physical
properties, explain the largest amount of variance in microbial
communities [1]. Soil pH impacts microbial and fungal diversity,
but temperature and latitude do not, and greater diversity is
found where soil pH is more basic [2]. Additionally, Arbuscular
mycorrhizal communities are dependent on pH, carbonate
content, base saturation, and phosphorus [3].
In addition to pH, nutrient availability impacts microbial
communities. Microbial biomass is directly related to soil organic
carbon content [4,5]. Not surprisingly, spatial distribution
of denitrifying organisms within a field is related to nitrates,
dissolved organic C, and dissolved organic N [6].
While pH and nutrients can individually impact distribution
and abundance of microbial communities, these factors can
also have a synergistic effect. Soil pH affects the availability
of nutrients. Wetter climates tend toward greater base cation
leaching, acidity, and fungi:bacteria ratios [7]. With less
moisture available for leaching, soil pH and base cation content
are typically greater.
Vegetation also impacts soil microbial communities. Globally,
microbial communities are impacted by biome and belowground
plant net primary production [8] while soil microbial community
composition varies with ecosystems [2]. For example, ratios of
fungi:bacteria are lower in grasslands due to differences in pH
and moisture. While both plants and soils impact soil microbial
communities [5], grassland type plays a major role in explaining
variability in soil microbial communities of grasslands [9]. When
considering both biotic and abiotic habitats, the predictive fit
of models in grazing pastures improves when plant traits are
included [7], and microbial communities can be quite plant-
specific [10]. Given that plants and soil microbes are intricately
related, we would expect that changing one would alter the
other. Microbial communities can be shaped by the presence of
species-specific root exudates, which can provide C and growth
factors for auxotrophic organisms [11].
In Texas, Ashe juniper (Juniperus ashei) has been overtaking
savannas previously dominated by live oak (Quercus virginiana).
This is a major management concern for water-limited Texas
Abstract
Soil microbial communities depend on pH, texture, and available nutrients. Ashe juniper (Juniperus ashei) is encroaching on live oak (Quercus
virginiana) savannas. Litter differences may alter soil microbial communities and therefore the ability of the soil ecosystem to cycle nutrients. The
objective was to determine whether soil microbial communities differ under the canopies of Ashe juniper and live oak. At each of three transects
we collected four surface soil samples under live oak canopy, overlapping canopies of live oak and Ashe juniper, under Ashe juniper canopy, and
in grass away from the trees. We characterized soils by pH, texture, calcium carbonate equivalence, total nitrogen, and total carbon. Microbial
communities were analyzed using massively parallel DNA sequencing and analyzed within QIIME using Kruskal-Wallis ANOVA, Mann-Whitney U,
and Spearman’s rank correlations. Prokaryotic communities differed along the transect, while fungal communities did not. Soil pH increased from
live oak to grass. Trends in taxa were mostly associated with vegetation, pH, and CCE. Many prokaryotic genera associated with soil properties
were important nutrient cyclers. Many fungal genera associated with pH and vegetation were mycorrhizal fungi, suggesting a plant-specific
relationship.
Keywords: 16S; ITS; Sequencing; Juniperus; Quercus; Vegetation; Rangeland; Soil health