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Original article doi: 10.1016/S2222-1808(15)60920-3 ©2015 by the Asian Pacific Journal of Tropical Disease. All rights reserved.
Acute toxicity of caprine alpha S2-casein protein on the microstructures and mineral profles of rat ileum
Vita Agustina
1
, Bambang Setiawan
2
, Fatchiyah Fatchiyah
1*
1
Department of Biology, Faculty of Mathematic and Natural Sciences, Brawijaya University, Malang 65145, East Java, Indonesia
2
Department of Medical Chemistry and Biochemistry, Medical Faculty, Lambung Mangkurat University, Banjarmasin, South Kalimantan, Indonesia
Asian Pac J Trop Dis 2015; 5(9): 721-725
Asian Pacific Journal of Tropical Disease
journal homepage: www.elsevier.com/locate/apjtd
*Corresponding author: Prof. Fatchiyah Fatchiyah, PhD, Department of Biology,
Faculty of Mathematic and Natural Sciences, Brawijaya University, Jl. Veteran,
Malang 65145, Indonesia.
Tel: +62341 575841
E-mail: fatchiya@ub.ac.id
Foundation Project: Supported by BOPTN-RU PTN UB decentralization research
grant of 2012-2014.
1. Introduction
Caprine casein protein contains mainly in αS1-, αS2-, β- and
κ-casein, of which those types of casein have different properties
and bioactive functions[1,2]. Bioactive peptides have potential as
antibacterial, immunomodulatory, antioxidant, antithrombotic and
antiinflammation activities[3-5]. Casein is potential as well to be
developed as a source of nutrients for the body. Casein derived a lot
of bioactive peptides that have a health benefit to control molecular
and/or cellular signaling mechanism. Recently in our study, we had
identified that the molecular weight of caprine αS2-casein (CSN1S2)
protein is separately electroporated at 36 kDa, in contrast we did not
find in bovine protein. The function of 36 kDa CSN1S2 protein had
been characterized as proper. This study reported that CSN1S2 (αS2)
protein isolated from Etawah goat milk consists of 223 amino acids
and 8 bioactive peptides, so that there are many interactive sites for
negatively and/or positively charged molecules[6]. Our previous
in vitro study reported that CSN1S2 affected the proliferation of
preosteblast cells[7]. Furthermore, in vivo study also supported the
inflammatory effect of this casein on complete Freund’s adjuvant
induced rheumatoid arthritis rats[8].
Ileum, the final part of the small intestine, performs the function
of absorbing nutrients which have been processed by the stomach.
The ileum consists of 4 layers: mucosa, sub-mucosa, muscularis
propria (oriented smooth muscles fibers) and outer serosa layer.
Mucosa epithelium folds to form villi that increase surface area
and sub-mucosa embodies glands that secrete digestive enzymes[9].
Observation using scanning electron microscope (SEM) showed that
mucosal villi are shaped like fingers which protrude into the lumen.
The villous surface is almost entirely covered with enterocytes
that act as absorptive cells[10]. Through absorption in the ileum,
milk proteins including the CSN1S2 proteins are broken down into
ARTICLE INFO ABSTRACT
Objective: To determine the microstructures and mineral profiles of ileum villi in rats treated
acutely with caprine αS2-casein (CSN1S2) protein isolated from Etawah goat milk.
Methods: Fifty (male and female) rats were divided into 5 groups, including a group of rats-
untreated as a control (untreated) and rats-treated with caprine CSN1S2 protein at doses of 500,
1 000, 2 000, and 4 000 mg/kg body weight by oral single dosage. The microstructures were
analyzed by scanning electron microscope. The mineral profiles of ileum were measured by
scanning electron microscope-energy dispersive X-ray software version 1.7. The degree of
ileum perforation was calculated by BoneJ software.
Results: The ileum microstructure of the control group showed a smooth and compact surface,
while the treating groups showed less compact surface and minimal perforation. The levels of
sodium, sulphur and phosphorus were statistically significant higher in treating females at all
doses compared to the control (P < 0.05). In contrast, the selenium levels were significantly
lower in treating females at all doses than that of the control group (P < 0.05).
Conclusions: This research concludes that the caprine CSN1S2 protein is able to stimulate the
ileum toxicity up to 2 000 mg/kg body weight dosage and change sodium, phosphorus, sulphur
and selenium homeostasis of ileum in female rats.
Contents lists available at ScienceDirect
Article history:
Received 25 May 2015
Received in revised form 26 May 2015
Accepted 1 Jun 2015
Available online 16 Jul 2015
Keywords:
Acute toxicity
Intestine
Functional food
Etawah goat milk