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J Endocrinol Invest
DOI 10.1007/s40618-016-0549-y
OPINION
Adipokines and bone metabolism: an interplay to untangle
J. Pepe
1
· C. Cipriani
1
· M. Cilli
1
· L. Colangelo
1
· S. Minisola
1
Received: 4 September 2016 / Accepted: 6 September 2016
© Italian Society of Endocrinology (SIE) 2016
is probably mediated by the leptin receptor found on osteo-
blasts and chondrocytes. The indirect effect is mediated
by different pathways. Leptin also impacts and regulates
osteocalcin, an osteoblast-derived hormone considered
a marker of bone formation, which in turn regulates both
bone metabolism and insulin sensitivity [4]. Serum leptin
levels have been shown to have a strong positive correla-
tion with bone mineral density (BMD) in women, and high
levels of leptin were reported to be predictive of low risk of
fractures in some but not all studies [4]. Leptin increases
osteoprotegerin (OPG), a member of the tumor necrosis
factor (TNF) alpha superfamily secreted by osteoblasts
which acts as a soluble decoy receptor for the activator of
nuclear factor kappa B (RANK), a transmembrane protein
expressed by osteoclasts. By preventing the soluble recep-
tor activator of nuclear factor kappa B ligand (RANKL)
from binding to RANK, OPG protects bone from excessive
resorption.
Adiponectin levels have been shown to be inversely
related to visceral fat and body mass index [5]; serum val-
ues vary largely depending also on age, sex, menopausal
status and comorbidities [6]. Secretion of adiponectin may
be inhibited by cytokines, such as TNF alpha, interleu-
kin 6 and hormones (cortisol and testosterone), which are
also involved in bone metabolism. Adiponectin receptors
have been demonstrated on osteoblasts, but also on mus-
cle and liver, which are organs that interplay with adipose
tissue and bone metabolism [7]. Osteoblasts and marrow
adipocytes originate from a common mesenchymal pro-
genitor, and adiponectin has been postulated as a poten-
tial factor that may induce osteoblast proliferation and
differentiation. Interestingly, osteocalcin may upregulate
the expression of the adiponectin gene in adipocytes, thus
improving insulin sensitivity. Adiponectin has been studied
also in different clinical conditions such as non-alcoholic
Recent evidence of increased fracture risk in obese patients
has focused the attention on the relationship between adi-
pose and skeletal tissue [1].
Adipocytes secrete a number of bioactive molecules,
such as adipokines, which recently have been shown to
have a role in bone metabolism. Adipokines are pleiotropic
molecules, investigated in many physiological or patholog-
ical processes, including inflammation, endothelial dam-
age, atherosclerosis, impaired insulin signaling and hyper-
tension. Adipokine dysregulation has been shown to be a
strong determinant of the low-grade inflammatory state of
obesity, which promotes a cascade of metabolic alterations
leading to cardiovascular complications, insulin resistance
or diabetes mellitus [2, 3]. So far, only a few adipokines
(adiponectin, leptin, resistin, visfatin and the newly discov-
ered omentin-1) have been studied in the context of bone
metabolism.
Leptin was the first adipokine discovered. Circulating
levels seem to reflect the amount of energy stored in the
adipose tissue; however, a wide variability in leptin levels
has been reported in individuals with the same body mass
index [4]. A number of factors regulate circulating leptin
levels in healthy individuals. In addition, leptin also influ-
ences several hypothalamic pituitary peripheral neuroen-
docrine axes, including the thyroid, gonadal, cortisol and
growth hormone axes.
Leptin exerts its effect on bone metabolism by means of
both direct and indirect mechanisms. The direct mechanism
* J. Pepe
jessica.pepe@uniroma1.it
1
Department of Internal Medicine and Medical Disciplines,
“Sapienza” University of Rome, Viale del Policlinico 155,
00161 Rome, Italy