Biotechnology and Bioprocess Engineering 2008, 13: 571-576
DOI/10.1007/s12257-008-0014-7
Effects of Inoculum Concentration,
Temperature, and Carbon Sources on
Tannase Production during Solid State
Fermentation of Cashew Apple Bagasse
=
Tigressa H. S. Rodrigues
1
, Gustavo A. S. Pinto
2
, and Luciana R. B. Gonçalves
1
*=
1
Universidade Federal do Ceará - Departamento de Engenharia Química, Campus do Pici, Bloco 709 - 60455-760, Fortaleza,
CE - Brazil
2
Embrapa Agroindústria Tropical, Rua Dra. Sara Mesquita, 2270, Planalto do Pici - 60511-110, Fortaleza, CE – Brazil=
Abstract In this study, the optimization of tannase production by solid state fermentation was investigated using cashew apple ba-
gasse (CAB), an inexpensive residue produced by the cashew apple agroindustry, as a substrate. To accomplish this, CAB
was enriched with 2.5% (w/w) tannic acid and 2.5% (w/w) ammonium sulphate and then moistened with water (60 mL/100 g
of dry CAB). The influence of inoculum concentration (10
4
to 10
7
spores/g), temperature (20, 25, 30, and 35°C) and several
additional carbon sources (glucose, starch, sucrose, maltose, analytical grade glycerol, and glycerol produced during bio-
diesel production) on enzyme production by ^ëéÉêÖáääìë= çêóò~É was then evaluated. Supplementation with maltose and
glycerol inhibited tannase synthesis, which resulted in lower enzyme activity. Starch and sucrose supplementation increased
enzyme production, but decreased the enzyme productivity. The maximum tannase activity (4.63 units/g of dry substrate)
was obtained at 30°C, using 10
7
spores/g and 1.0% (w/v) sucrose as an additional carbon source. © KSBB
hÉóïçêÇëW=Å~ëÜÉï=~ééäÉ=Ä~Ö~ëëÉI=ëçäáÇ=ëí~íÉ=ÑÉêãÉåí~íáçåI= Aspergillus oryzaeI=í~åå~ëÉ=
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INTRODUCTION
Solid state fermentation (SSF) is a bioprocess in which
microorganisms are grown on solid substances in the ab-
sence or near absence of free water [1]. In general, SSF takes
place under low water/high solids conditions [2]. Two types
of substrates are used in SSF, those that are used by the mi-
croorganisms as the carbon and energy source, and those that
only support the fermentation process [3]. Due to its low cost
and effectiveness, SSF is often the system of choice for con-
verting agro industrial residues to fermented foods, enzymes,
and other extracellular metabolites such as organic acids and
flavour compounds [2].
The cashew apple is a pseudofruit native to Brazil that is
primarily grown in the North and Northeast regions of Brazil,
where the cashew agroindustry plays a substantial role in the
local economy. Cashew apples are hard, pear-shaped, small,
*Corresponding author
Tel: +55-85-3366-9611 Fax: +55-85-3366-9610
e-mail: lrg@ufc.br
nonclimacteric fruit that occur in three colours: yellow, or-
ange, and red. The primary product of the cashew apple is
the cashew nut, which is well known around the world. The
edible portion, which comprises 90% of the fruit, is a
pseudofruit rich in vitamin C, flavour, and aroma. However,
the industrially processed pseudofruit products are consumed
by the local market and do not play an important role in the
state or national economy [4-7].
When the pseudofruit is industrially processed for the pro-
duction of juice, 40% (w/w) of the fruit remains as bagasse.
This bagasse is not used for human consumption and is gen-
erally discarded. However, due to its composition (Table 1),
cashew apple bagasse (CAB) represents an interesting and
inexpensive (less than US$ 0.50/kg) substrate for several
applications, including the production of microbial enzymes
by solid state fermentation (SSF) [8-10].
Tannase (tannin acyl hydrolase, E.C. 3.1.1.20) catalyzes
the hydrolysis of bonds present in the molecules of tannins
and gallic acid esters [11]. Tannase is applied commercially
in the food and drink industries, as well as in the production
of pharmaceutically important compounds such as gallic acid.