Protein Quality and Antinutritional Factors of Wild Legume Seeds
from the Sonoran Desert
Magdalena Ortega-Nieblas*
Center of Scientific and Technologic Research, University of Sonora (CICTUS), A.P. 1819,
Hermosillo, Sonora, Mexico 83000
Luz Va ´ zquez-Moreno and Marı ´a R. Robles-Burguen ˜o
Research Center for Food and Development (CIAD), A.P. 1735, Hermosillo, Sonora, Mexico 83000
Seeds from seven species of legume trees of the Sonoran Desert were studied to determine their
potential value as a nonconventional food and/or as a feed source. Seed proximal composition and
protein quality were evaluated by conventional methodology. Defatted flours from Acacia farnesiana,
Cercidium microphyllium, Cercidium sonorae, Mimosa grahamii, Olneya tesota, Parkinsonia
aculeata, and Prosopis juliflora have protein contents that ranged from 20 to 30%. These proteins
were deficient in sulfur amino acids but high in lysine and phenylalanine content. Except for M.
grahamii seeds, trypsin inhibitors were relatively high in all species and varied from 50 to 70 TIU/
g. Other antinutritional factors as lectins, alkaloids, saponins, and phenols were detected in all
but Mimosa seeds. Protein in vitro digestibility was found to be 77-84%, and these values increased
(5-10%) after a thermic treatment. The calculated C-PER values were found to be 1.8-2.1%. From
the seeds of the seven species evaluated, M. grahamii showed the highest potential as a protein
source.
Keywords: Desert legume seeds; protein quality; antinutritional factors
INTRODUCTION
Intense efforts to find alternative sources of proteins
from plants adapted to adverse conditions are been
conducted around the world (Siddhuraju et al., 1995;
Bravo et al., 1994; Bhattacharya et al., 1994). Seeds
from Cassia laevigata and Tamaridus indica, both
legumes from India, were reported as a low-cost protein
source to alleviate the protein malnutrition among
people living in developing countries (Siddhuraju et al.,
1995). Seeds and pods from the genera of Acacia,
Olneya (palo fierro, iron wood), and Prosopis (mesquite)
have been used in several countries for human con-
sumption, livestock food, and fodder production (Figueire-
do, 1990; Degen et al., 1995; Felker, 1981). Studies of
mesquites from South America and the Mediterranean
confirmed that these plants produce pods and seeds that
are palatable to human and animals. In a recent
comparison of mesquite pods with carob pods (the latter
is widely used in the food industry), it was demonstrated
that mesquite pods had better nutritional properties
than carob pods, including lower polyphenolic content
and higher protein digestibility (Bravo et al., 1994).
In Mexico, the Sonoran Desert has a large group of
perennial trees from the Leguminosae family that
produce large amounts of indehiscent pods; however, the
chemical composition and protein quality of their seeds
remain unknown. These plants have important agro-
nomic characteristics applicable under unfavorable
conditions such as reduced availability of water and
fertilizers (Felker, 1981; Meyer et al., 1986; Bresanni
and Elias, 1974). In the United States, some of these
species have been used for arid land reforestation and
erosion control (Estrada and Marroquin, 1992). Fur-
thermore, Prosopis pod yield has been estimated up to
10 000 kg/ha (Felker, 1981; Felger and Nabham, 1978).
In this work, seeds of seven legume species of the Sonora
Desert were selected based on their abundance in the
natural habitat and productivity, and their seed com-
position and protein quality were determined by con-
ventional methods.
MATERIALS AND METHODS
Seeds from selected species were collected from various
regions of the Sonoran Desert (Rodney et al., 1972; Ortega-
Nieblas, 1993). The species were Acacia farnesiana (huizache),
Cercidium microphyllium (palo verde 1), Cercidium sonorae
(brea), Mimosa grahamii (gatun ˜ a), Olneya tesota (palo fierro,
iron wood) Parkinsonia aculeata (palo verde 2), and Prosopis
juliflora (mesquite). Seeds were manually separated from
husks and ground on a Wiley grinder using a 150 μm mesh.
All reagents used were of analytical grade.
Chemical Analysis. Fat was extracted from 100 g of meal
in a Soxhlet apparatus using hexane (AOAC, 1990). Defatted
flours were stored at 4 °C until used. The nitrogen content of
flours was determined using the micro-Kjeldahl method
(AOAC, 1990). A conversion factor of 6.25 was used to
calculate protein content. Moisture, crude fiber, and ash were
determined using the official methods (AOAC, 1990). Carbo-
hydrate content was calculated by difference.
Antinutritional Factors. Trypsin inhibitors were deter-
mined by using the procedure described by Hamerstrand and
Black (1981). Phenols, alkaloids, and saponins were deter-
mined using the methods described by Goldstein and Swain
(1975), Waldi and Schnackerz (1973), and Harbone (1984).
Hemagglutinin Activity. Crude extracts were obtained
by extraction of flours in saline solution, 1:10 (w/v), for 2 h at
room temperature and overnight at 4 °C. Crude extracts were
recovered after centrifugation, 20 min at 5000g (Lis and
Sharon, 1986). Hemagglutinin activity was determined by the
serial double-dilution method using human B
+
erythrocytes
treated with trypsin (Turner and Liener, 1975).
* Author to whom correspondence should be ad-
dressed [fax (62) 80-0058].
3130 J. Agric. Food Chem. 1996, 44, 3130-3132
S0021-8561(95)00613-3 CCC: $12.00 © 1996 American Chemical Society