Vol.:(0123456789) 1 3
Environmental Sustainability
https://doi.org/10.1007/s42398-019-00055-3
ORIGINAL ARTICLE
Cadmium resistant plant growth promoting rhizobacteria Serratia
marcescens S2I7 associated with the growth promotion of rice plant
Rhitu Kotoky
1
· Sudipta Nath
1
· Dinesh Kumar Maheshwari
2
· Piyush Pandey
1
Received: 17 December 2018 / Revised: 16 April 2019 / Accepted: 4 May 2019
© Society for Environmental Sustainability 2019
Abstract
Cadmium (Cd) is one of the most toxic contaminants in the environment. The contamination of crop felds with metals like
Cd may lead to a signifcant decrease in productivity due to related toxicity. The present study was conducted with aim to
enhance the growth of rice (Oryza sativa), in Cd contaminated soil, by the application of heavy metal resistant—plant growth
promoting rhizobacteria, Serratia marcescens S2I7. S2I7 was found to be highly resistant to Cd and positive for phosphate
solubilization, production of siderophore, indole acetic acid (IAA), and hydrogen cyanide (HCN). The organism was found
to have Glutathione S-transferase (GST) mechanism for detoxifcation of Cd. GST was induced and its activity was increased
signifcantly in the presence of Cd. When the bacterial strain was applied to rice seedlings, the germination index and growth
of seedling were improved in spite of Cd contamination in soil. Pot trial experiments in laboratory conditions showed that
the growth of the plants was not afected up to the moderate concentration of Cd (0.75 M), and removal of Cd from the soil
was also high (66 mg/kg after 20 days). Interestingly, at low concentration of Cd (0.25 M), there was no signifcant removal
of residual Cd from the soil. However, under the high concentration, rate of removal of Cd was very less after 20 days which
may be due to toxic efects (of higher concentration). Here the plant–microbe association may need more time to overcome
the efects of Cd and to lead to removal of the metal. Moreover, with the application of isolate S2I7, the growth of the rice
plant was comparatively better, as the mean shoot length was increased by 7.9%, and mean root length was increased by
53.7%. Therefore, the application of multifunctional plant-growth promoting bacteria exhibiting resistance for Cd may result
in better growth of rice under the stress of the heavy metal. This may also improve the remediation of contaminated sites by
alleviating Cd-induced phytotoxicity and promoting the growth of plants.
Keywords Cadmium · PGPR · Oryza sativa · Bioremediation Serratia marcescens
Introduction
Increasing industrialization has led to heavy metal contami-
nation of the environment, which has posed various threats
to humans, other living organisms and the stability of the
ecosystems. Unlike organic contaminants, heavy metals
are not biologically degradable, and therefore remain in the
environment for a long time. Most of the heavy metals are
categorized as toxic (Wood 1974), but the toxicity of difers
greatly with the metal (Fergusson 1990). Cadmium (Cd) and
lead (Pb) cause toxicity to microorganisms by binding to
essential respiratory proteins (Vallee and Ulmer 1972), and
through oxidative damage by the production of reactive oxy-
gen species (ROS) (Stohs and Bagchi 1995). As Cd and Pb
are currently not known to have any biological function in
bacteria therefore, less concentration of these may also dis-
turb the normal functioning of the microorganisms (Kotoky
and Pandey 2018).
Microorganisms play an important role in the biogeo-
chemical cycling of heavy metals and therefore in cleaning
up metal-contaminated soil. Microorganisms, exposed to the
higher concentrations of toxic heavy metals develop resist-
ance against the elevated levels of these metals. Such metal
resistant microorganisms are useful for successful bioreme-
diation, as they use diferent detoxifcation mechanisms that
are helpful to remove the contaminants. Apart from this,
microorganisms play a crucial part in the growth of plants
* Piyush Pandey
piyushddn@gmail.com
1
Department of Microbiology, Assam University, Silchar,
Assam 788011, India
2
Department of Botany & Microbiology, Gurukul Kangri
University, Haridwar 249404, Uttarakhand, India