J Appl Ichthyol. 2017;1–3. wileyonlinelibrary.com/journal/jai
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1 © 2017 Blackwell Verlag GmbH
Received: 1 December 2016
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Accepted: 28 March 2017
DOI: 10.1111/jai.13413
SHORT COMMUNICATION
New records of Speleogobius trigloides Zander & Jelinek, 1976
and S. llorisi Kovačić, Ordines & Schliewen, 2016 (Osteichthyes:
Gobiidae) in the Aegean Sea
S. Engin | A. Ç. Oruç | D. Seyhan | E. Irmak
Faculty of Fisheries, Izmir Katip Celebi University, Izmir, Turkey
Correspondence
Semih Engin, Faculty of Fisheries, Izmir Katip Celebi University, Izmir, Turkey.
Email: engin.semih@gmail.com
1 | INTRODUCTION
The family Gobiidae is the largest fish family in the Turkish ma-
rine waters with 48 species (9.2% of the total marine ichthyofauna)
(Bilecenoğlu, Kaya, Cihangir, & Çiçek, 2014; Engin, Irmak, Seyhan,
Akdemir, & Keskin, 2016). Recently, 33 gobiid fish species were veri-
fied in the north-eastern area of the Aegean Sea (Engin et al., 2016).
The genus Speleogobius is represented by two species: Speleogobius
trigloides Zander & Jelinek, 1976 and the recently described
Speleogobius llorisi Kovačić, Ordines and Schliewen, 2016. The pres-
ent study represents new information on the range extension and
morphological features of the Speleogobius species reported from the
Aegean Sea.
2 | MATERIAL AND METHODS
The underwater surveys were conducted with two closed-circuit
rebreather (eCCR) divers. One female specimen of S. trigloides sam-
pled from Melina Island/Ayvalık (39°20′08.34″N; 26°33′36.73″E;
Figure 1) at 33 m depth in June 2016 and one male specimen of S. llorisi
sampled from Mardaliç Island (38°54′39.67″N; 26°49′28.50″E;
Figure 1) (Çandarlı) at 43 m depth in July 2016. The S. trigloides was
captured barehanded and fixed 70% alcohol; the S. llorisi was sampled
by hand net and fixed in 4% formaldehyde solution. Both specimens
were deposited in the Marine Life Collection Centre of İzmir Katip
Celebi University, Turkey (IKC.PIS.1090-91).
The specimens were identified according to Miller (1986) and
Kovačić, Ordines, and Schliewen (2016). Terminology of the lateral line
system followed Sanzo (1911), Miller (1986), and Kovačić et al. (2016).
Specimen measurements were taken under a stereomicroscope using
a digital calliper point-to-point (accuracy of 0.01 mm) within 2 days
after fixation. Scales in lateral series were counted from the axilla
along the lateral midline, including scales over the origin of C. In the
counts D2 and A, the last bifid ray was counted as one. Body depth
was measured at anal fin origin, caudal peduncle length measured at
end of A base to origin of C, head length measured from snout to mid-
line opposite upper origin of opercle, head width measured between
upper origin of post opercle, head depth measured between upper
origin of posterior part of eyes.
3 | RESULTS
Speleogobius specimens from the Aegean Sea were distinguishable
from other Gobiidae families in the Mediterranean Sea: pelvic fins
united into a disc; anterior membrane absent and posteriorly emargin-
ated; suborbital sensory papillae row present; posterior oculascapular
head canal absent; predorsal area scaled.
Speleogobius trigloides and S. llorisi are separated from each other
by several morphological characters: preopercular canal absent vs.
preopercular canal present with pores γ, δ, ε; short snout vs. long
snout; upper lip slightly protruding over lower lip vs. lower lip slightly
protruding over upper lip; scale in transversal series 6 vs. 7. Detailed
morphometric measurements are given in Table 1. First dorsal S. tri-
gloides fin has red-brown spots. There are brownish and vertical bands
on the trunk of S. llorisi.
The grotto goby S. trigloides has a moderately-elongated body and
a slender caudal peduncle. Head acute. Snout short, upper lip slightly
protruding over lower lip. Large eyes. Predorsal area covered with
seven rows of cycloid scales. Preopercular head canal absent.
Lively coloration. General body reddish, with five pale saddles on
the dorsal part of the trunk and eight pale blotches on the ventral part.
Caudal fin transparent-yellowish, dorsal fins redish-brown with violet