Journal Pre-proof Dinoflagellates encystment with emphasis on blooms in Boughrara Lagoon (South- Western Mediterranean): Combined effects of trace metal concentration and environmental context Fatma Abdmouleh Keskes, Najla Ayadi, Abdelfattah Atoui, Mabrouka Mahfoudi, Moufida Abdennadher, Lamia Dammak Walha, Sana ben Ismail, Olfa ben Abdallah, Yosra Khammeri, Marc Pagano, Asma Hamza, Malika Belhassen PII: S0272-7714(19)30722-X DOI: https://doi.org/10.1016/j.ecss.2020.106648 Reference: YECSS 106648 To appear in: Estuarine, Coastal and Shelf Science Received Date: 22 July 2019 Revised Date: 21 December 2019 Accepted Date: 13 February 2020 Please cite this article as: Abdmouleh Keskes, F., Ayadi, N., Atoui, A., Mahfoudi, M., Abdennadher, M., Walha, L.D., Ismail, S.b., Abdallah, O.b., Khammeri, Y., Pagano, M., Hamza, A., Belhassen, M., Dinoflagellates encystment with emphasis on blooms in Boughrara Lagoon (South-Western Mediterranean): Combined effects of trace metal concentration and environmental context, Estuarine, Coastal and Shelf Science (2020), doi: https://doi.org/10.1016/j.ecss.2020.106648. This is a PDF file of an article that has undergone enhancements after acceptance, such as the addition of a cover page and metadata, and formatting for readability, but it is not yet the definitive version of record. This version will undergo additional copyediting, typesetting and review before it is published in its final form, but we are providing this version to give early visibility of the article. Please note that, during the production process, errors may be discovered which could affect the content, and all legal disclaimers that apply to the journal pertain. © 2020 Published by Elsevier Ltd.