1 This article is protected by copyright. All rights reserved. Rice cyclophilin OsCYP18-2 is translocated to the nucleus by an interaction with SKIP and enhances drought tolerance in rice and Arabidopsis 1 Running title: SKIP interacting cyclophilin enhances drought tolerance Sang Sook Lee 1, ǂ , Hyun Ji Park 1, ǂ , Dae Hwa Yoon 1, 2 , Beom-Gi Kim 3 , Jun Cheul Ahn 1, 2 , Sheng Luan 4, *, Hye Sun Cho 1, * 1 Sustainable Bioresource Research Center, Korea Research Institute of Bioscience and Biotechnology, Daejeon 305-806, Korea, 2 Department of Pharmacology, College of Medicine, Seonam University, Namwon 590-170, Korea, 3 Divisions of Bio-Crops Development, National Academy of Agricultural Science, RDA, 441-707, Korea, 4 Department of Plant and Microbial Biology, University of California at Berkeley, Berkeley CA 73072, USA ǂ These authors contributed equally to this work *Co-corresponding authors: S. Luan e-mail address: sluan@berkeley.edu H. S. Cho e-mail address: hscho@kribb.re.kr This article has been accepted for publication and undergone full peer review but has not been through the copyediting, typesetting, pagination and proofreading process, which may lead to differences between this version and the Version of Record. Please cite this article as doi: 10.1111/pce.12531 Accepted Article