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Eur Food Res Technol
DOI 10.1007/s00217-015-2442-x
ORIGINAL PAPER
A large diversity of lactic acid bacteria species is involved
in the fermentation of wheat used for the manufacture of lemzeiet
Ryma Merabti · Farida Bekhouche · Victoria Chuat ·
Marie N. Madec · Marie B. Maillard · Sylviane Bailly ·
Anne Thierry · Florence Valence
Received: 1 December 2014 / Revised: 8 February 2015 / Accepted: 13 February 2015
© Springer-Verlag Berlin Heidelberg 2015
very similar profiles for both processes. Sixty-nine isolates
were identified as belonging to six genera of 16 species
(Enterococcus, Lactobacillus, Leuconostoc, Pediococ-
cus, Weissella, and Streptococcus). The profiles of volatile
aroma compounds showed a marked effect of the fermenta-
tion process, compared to non-fermented wheat, with 35 of
the 40 volatiles detected at amounts 20- to 30-fold higher in
fermented wheat samples. This study gives the first insight
into lactic acid bacteria population diversity and activity in
fermented wheat and will contribute to a better control of
the fermentation process.
Keywords Lactic acid bacteria · Fermentation · Cereal ·
Vinegar · Volatile
Abbreviations
ANOVA Analysis of variance
BHYE Brain heart infusion yeast extract
CFU Colony forming unit
DMS Dimethyl sulfide
DMDS Dimethyl disulfide
FA Fat acidity
GC–MS Gas chromatography–mass spectrometry
HS Headspace
LAB Lactic acid bacteria
LSD Least significant difference
MA Malt agar
MRS Man Rogosa and Sharpe
OAIC Office algérien interprofessionnel des céréales
PCA Principal component analysis
pheS Phenylalanyl-t-RNA synthetase alpha-subunit
PFGE Pulsed-field gel electrophoresis
RAPD Random amplified polymorphic DNA
rpoA RNA polymerase alpha-subunit
TTA Titratable acidity
Abstract Algerian couscous named lemzeiet is manu-
factured from fermented wheat. Historically performed
in underground silos called matmor, the fermentation of
wheat is now generally carried out in plastic jerrycans with
or without addition of vinegar at the beginning of the fer-
mentation. Culture-dependent and culture-independent
methods (PCR-TTGE) were used to characterize lactic acid
bacteria and to determine their dynamic and diversity over
a two-year period, with and without the addition of vinegar.
Fungi, physicochemical characteristics, and volatile com-
pound profiles were also monitored. The isolates obtained
from different stages of fermentation and from both pro-
cesses were characterized by coupling different molecular
methods (16SrRNA/pheS/rpoA gene sequencing, species-
specific PCR, RAPD and PFGE). PCR-TTGE revealed
R. Merabti (*) · F. Bekhouche
Département de Biotechnologie Alimentaire, Institut
de la Nutrition, de l’Alimentation et des Technologies
Agro-alimentaires (INATAA), Université Constantine 1,
25000 Constantine, Algeria
e-mail: merabtiryma@yahoo.fr
R. Merabti · V. Chuat · M. N. Madec · M. B. Maillard ·
A. Thierry · F. Valence (*)
UMR1253, Science et Technologie du Lait et de l’Œuf,
CIRM-BIA, INRA, 65 rue de Saint Brieuc, 35000 Rennes,
France
e-mail: Florence.Valence-Bertel@rennes.inra.fr
V. Chuat · M. N. Madec · M. B. Maillard · A. Thierry · F. Valence
UMR1253, Science et Technologie du Lait et de l’Œuf,
Agrocampus Ouest, 65 rue de Saint Brieuc, 35000 Rennes,
France
S. Bailly
UMR1331, Toxalim, Research Centre in Food Toxicology, INRA,
31027 Toulouse, France