Brain Research, 338 (1985) 341- 345 341
Elsevier
BRE 20924
Retinal and tectal responses to alternating gratings are
unaffected by monocular deprivation in pigeons
PAOLA BAGNOLP, VITTORIO PORCIATTI 2and WALTER FRANCESCONI 1
llstituto di Fisiologia dell'Universitd e lstituto di Neurofisiologia del C.N.R., 56100 Pisa and
2Divisione Oculistica, USL 13, 57100 Livorno (Italy)
(Accepted February 19th, 1985)
Key words: electroretinogram - - tectal evoked potential - - grating - - monocular deprivation - - pigeon
Retinal and tectal potentials to alternating gratings were recorded in pigeons raised monocularly deprived. Deprived eyes showed
consistent myopia and anterior-posterior axis elongation. In addition, vitreal opacities were observed in 70% of the deprived eyes. In
pigeons with vitreal opacities, a reduction of retinal and tectal response amplitude was consistently found in the high spatial frequency
range. Pigeons with clear media of the deprived eyes showed normal retinal and tectal responses.
Although the morphofunctional changes observed
in the mammalian geniculocortical pathway after
prolonged early monocular deprivation are extreme-
ly dramatic and well documented, questions remain
concerning the extent of the deprivation-induced
changes at the retinal level.
On the one hand, kittens either monocularly de-
prived or raised with a convergent squint showed a
reduction in the spatial resolving power of retinal
ganglion cells6:3: 9. Reduction in pattern-electroreti-
nogram (ERG) amplitude was also shown in human
amblyopes~,25, and a reduction in the ERG b-wave
was reported in dark-reared kittens S and guinea
pigs is. In addition, morphological and biochemical
changes were reported at the retinal level in animals
exposed to varying visual environmentslS,21.26,28 (see
ref. 3 for references).
On the other hand, several studies on monocularly
lid-sutured kittens have consistently reported normal
spatial resolution7,J7 and normal receptive field prop-
erties24 of deprived 'X' and 'Y' ganglion cells.
Recently, the avian visual system has provided a
useful model in searching for the neuronal correlates
of early visual deprivation. A dramatic loss of binoc-
ularly driven neurons was demonstrated in the visual
Wulst of owls raised with one eye sutured 20. In addi-
tion, significant deficits in pattern discrimination
learning and interocular transfer were described in
early monocularly deprived pigeons 5. Furthermore,
preliminary results have indicated a behavioral im-
pairment of the spatial acuity of pigeons' deprived
eyes (Burkhalter and Kniisel, personal communica-
tion).
The aim of this work was to determine whether
pattern-ERG and tectal surface-evoked potentials
(TEPs) were affected by early monocular depriva-
tion.
In monocularly deprived pigeons, we compared
the amplitudes of ERG and TEP to alternating grat-
ings of various spatial frequencies presented to the
deprived eyes to those of the responses elicited by the
stimulation of the undeprived eyes.
Experiments were performed on 10 pigeons of
both sexes weighing 350-400 g which were monocu-
larly deprived during the first 6 months of life by cov-
ering one eye with a black plastic cap immediately af-
ter hatching (for details of the procedure see Burk-
halter and Cu6nod5).
Details of the electrophysiological technique have
been previously describedL Briefly, the pigeons
Correspondence: P. Bagnoli, Instituto di Fisiologia dell'Universit~ e Istituto di Neurofisiologia del C.N.R., Via S. Zeno, 31-51,
56100 Pisa, Italy.
0006-8993/85/$03.30 © 1985 Elsevier Science Publishers B.V. (Biomedical Division)