ISSN 0006-3509, Biophysics, 2011, Vol. 56, No. 6, pp. 1069–1073. © Pleiades Publishing, Inc., 2011.
Original Russian Text © G.V. Maksimov, N.A. Brazhe, A.I. Yusipovich, E.Yu. Parshina, O.V. Rodnenkov, A.B. Rubin, G.G. Levin, V.A. Bykov, 2011, published in Biofizika, 2011,
Vol. 56, No. 6, pp. 1099–1104.
1069
INTRODUCTION
A topical task in nanobiotechnology is to design
integrated methods of cell microscopy that would
assess not only the morphology of subcellular nano-
structures but also their functional state. Here we dis-
cuss the merits and specific features of integrating
atomic force microscopy (AFM), laser interference
microscopy (LIM), and Raman scattering microscopy
(RSM) in erythrocyte studies.
AFM is used to determine the geometric dimen-
sions and relief of the cell. LIM allows examining low-
contrast biological objects without using dyes; to add,
knowing the refractive index, one can thereby deter-
mine the content and distribution of substance in the
cell [1]. With RSM, it is possible to assess not only the
distribution of but also the conformational changes in
the molecules of interest by the features of Raman
spectra.
Each of these methods separately is also used to
help detect various pathological states. The shape and
volume of fixed erythrocytes estimated by AFM [2] is
a diagnostic trait for a number of pathologies [3]. LIM
has revealed a significant increase in erythrocyte area
and volume in patients with ischemic heart disease [4],
which appears to be connected with a change in
plasma membrane viscosity and charge, as well as with
water accumulation in the erythrocyte because of acti-
vated Na/H exchange.
Altered conformation of hemoglobin (Hb)
revealed by RS in cardiovascular patients [5] and in
experimental rat brain ischemia [6] is supposed to
result from changes in the membrane characteristics
and redistribution of Hb in erythrocytes.
Combination of AFM, LIM and RSM would thus
permit assessing the change in geometry and redistri-
bution of cytoplasmic compartments as well as the dis-
tribution and conformation of Hb hemoporphyrin.
Visualization of the distribution of organelles and
plasma proteins, Hb in particular, provides valuable
information on cell functioning, which can be used for
studying the mechanisms of oxygen transport by
erythrocytes and fine interactions among cell
organelles. To add, joint use of these methods allows
substantial reduction of the required sample volume.
The erythrocyte is a widespread object in biomedi-
cal research. Many drugs are introduced into the
bloodstream, so it is important to be able to analyze
their impact of erythrocyte Hb. It is now customary to
distinguish two fractions: free cytosolic Hb
c
and mem-
brane-associated Hb
m
; the latter makes ~0.5% of the
total. In present-day notions, Hb
m
interacts with the
cytosolic part of the AE1 transporter (band 3 protein),
as does ankyrin of the submembrane cytoskeleton
(Fig. 1).
Changes in the properties of the plasma membrane
affect the Hb
m
conformation. Therefore, it is supposed
that early erythrocyte pathologies may be detected by
conformational alterations in submembrane Hb. It is
such nanoscale alterations in Hb
m
that are revealed by
surface-enhanced Raman scattering (SERS) [7],
which is an excellent supplement to AFM и LIM.
CELL BIOPHYSICS
Use of Nanoparticles for Studying the Conformations
of Submembrane Hemoglobin
G. V. Maksimov
a
, N. A. Brazhe
a
, A. I. Yusipovich
a
, E. Yu. Parshina
a
, O. V. Rodnenkov
b
, A. B. Rubin
a
,
G. G. Levin
c
, and V. A. Bykov
d
a
Biological Faculty, Moscow State University, Moscow, 119991 Russia
b
Russian Cardiology Research-and-Practice Complex, Moscow, 121552 Russia
c
All-Russia Research Institute for Optical and Physical Measurements, Moscow, 119361 Russia
d
NT-MDT Co., Zelenograd, Moscow 124482 Russia
Received June 9, 2011
Abstract—The advantages and specific features of integrated application of atomic force microscopy, laser
interference microscopy, and Raman microscopy in the study of erythrocytes are discussed. For successful
application of Raman microscopy in the surface-enhanced mode, use was made of silver colloids. The depen-
dence of the enhancement of Raman signals on silver nanoparticle size is demonstrated. The use of developed
methods in clinical diagnostics is discussed.
Keywords: erythrocyte, hemoglobin, Raman spectroscopy, silver colloids, diagnostics.
DOI: 10.1134/S0006350911060121