ISSN 1068-1620, Russian Journal of Bioorganic Chemistry, 2013, Vol. 39, No. 2, pp. 113–120. © Pleiades Publishing, Ltd., 2013.
Original Russian Text © A.V. Kamynina, M.P. Filatova, D.O. Koroev, A.Y. Abramov, O.M. Volpina, 2013, published in Bioorganicheskaya Khimiya, 2013, Vol. 39, No. 2, pp. 131–140.
113
1 2
INTRODUCTION
One of the hypothesis of Alzheimer’s disease devel-
opment involves accumulation of pathogenic form of
β-amyloid peptide (1-40) or (1-42), its further aggre-
gation leading to cell death and memory dysfunction.
Alzheimer’s disease is associated with such molecular
abnormalities in brain cells as oxidative stress and pro-
duction of reactive oxygen species (ROS) [1], alter-
ations in Ca
2+
-homeostasis (Ca
2+
-signaling) in neu-
rons and glial cells [2, 3], microglia activation and
other disturbances resulting in cell death. Prion pro-
tein was shown to accumulate in β-amyloid plaques
[4]. Detailed investigation revealed that soluble oligo-
mers of β-amyloid bind prion protein leading to dis-
turbances in long term potentiation in neurons and
Alzheimer’s disease development in transgenic mice.
Fragment 95-110 of the prion (PrP-(95-110)) was
shown to play an important role in its binding to
β-amyloid [5].
Monoclonal antibodies against fragment PrP-(95-
110) effectively restored memory dysfunctions in
Abbreviations: Aβ, β-amyloid; AEBSF, 4-(2-Aminoethyl) ben-
zenesulfonyl fluoride; Ahx, ε-aminocaproic acid; CFA, com-
plete Freund adjuvant; IFA, incomplete Freund adjuvant; PBS,
buffer containing 137 mM NaCl, 2.7 mM KCl, 7.8 mM
Na
2
HPO
4
, 1.5 mM KH
2
PO
4
, pH 7.4; PrP, prion protein; ROS,
reactive oxygen species.
1
The article was translated by the authors.
2
Corresponding author: phone: 8 (495) 336-57-77; e-mail:
aneskaminina@mail.ru.
transgenic mice and prevented β-amyloid binding
with prion protein in vivo [6]. However, the experi-
ments did not explore the mechanism of the protective
effect of the antibodies. In our work we investigate an
effect of monospecific affinity purified antibodies
against fragment PrP-(95-123) (PrP-(95-123)-MA)
in rat brain primary cultures of neurons and astrocytes
treated with β-amyloid. We earlier showed that anti-
bodies against PrP-(95-123), an extended analogue of
PrP-(95-110), bound both a recombinant prion and
native prion proten and were effective in preventing
prion disease development on cell lines [7]. The aim of
the current work is investigating an effect of PrP-(95-
123)-MA against β-amyloid toxi-city in cultured neu-
rons and astrocytes of cortex and hippocampus of rat
brain.
RESULTS AND DISCUSSION
Synthetic peptides and blood sera obtaining. Pep-
tide 95-123 of bovine prion protein was used for
immunization of animals (Table 1). Peptide construc-
tion PrP-(95-110)-TT combining rat prion fragment
95-110 and tetanus toxoid (TT) protein fragment 830-
844 [8] through ε-aminohexane acid was used for
investigating antibody binding in ELISA (Table 1).
The peptides were synthesized using the solid-phase
Fmoc-chemistry and characterized by HPLC and
mass spectrometry.
Antibodies to Synthetic Fragment 95-123 of the Prion Protein
Protect Neurons and Astrocytes from Beta-Amyloid Toxicity
1
A. V. Kamynina
a, 2
, M. P. Filatova
a
, D. O. Koroev
a
, A. Y. Abramov
b
, and O. M. Volpina
a
a
Shemyakin–Ovchinnikov Institute of Bioorganic Chemistry, Russian Academy of Sciences,
ul. Miklukho-Maklaya 16/10, Moscow, 117997 Russia
b
Institute of Neurology University College London, Queen Square, London WC1N 3BG, UK
Received August 20, 2012; in final form, October 16, 2012
Abstract—Molecular mechanisms of β-amyloid toxic effect on brain cells during Alzheimer’s disease is asso-
ciated with oxidative stress, intracellular Ca
2+
increase in neurons and astrocytes and activation of reactive
oxygen species production. Prion protein is known to be involved in beta-amyloid toxicity. Here we investi-
gated an effect of affinity purified antibodies to synthetic fragment 95-123 of the prion protein (PrP-(95-
123)) on β-amyloid induced cell death, Ca
2+
-signal, reactive oxygen species production and caspase 3 acti-
vation. We have shown that antibodies to PrP-(95-123) are able to protect neurons and astrocytes from beta-
amyloid induced cell death with no effect on the intracellular Ca
2+
concentration altered by beta-amyloid
treatment. However, the antibodies significantly reduced β-amyloid induced reactive oxygen species produc-
tion in astrocytes and decreased caspase 3 activation in neurons and astrocytes. Thus, induction of antibodies
to PrP-(95-123) of the prion protein provides a new approach to anti-Alzheimer’s disease vaccine design.
Keywords: synthetic peptide, prion protein, Alzheimer’s disease, cell culture, reactive oxygen species
DOI: 10.1134/S1068162013020076