1 Original article 1 2 Unique genetic features of canine adenovirus type 1 (CAdV-1) infecting red foxes (Vulpes vulpes) in Northern 3 Norway and arctic foxes (Vulpes lagopus) in Svalbard 4 5 Andrea Balboni 1 *, Morten Tryland 2 , Torill Mørk 3 , Siw T. Killengreen 4 , Eva Fuglei 5 , Mara Battilani 1 . 6 7 1 Department of Veterinary Medical Sciences, Alma Mater Studiorum-University of Bologna, Via Tolara di Sopra 50, 8 40064 Ozzano Emilia (BO), Italy. 9 2 Arctic Infection Biology, Department of Arctic and Marine Biology, UiT Arctic University of Norway, Framstredet 10 39, NO-9037 Tromsø, Norway. 11 3 Section of Pathology Oslo and Tromsø, Department of Analysis and Diagnostics, Norwegian Veterinary Institute, 12 Stakkevollveien 23, NO-9010 Tromsø, Norway. 13 4 Department of Education, UiT Arctic University of Norway, NO-9010 Tromsø, Norway. 14 5 Norwegian Polar Institute, Fram Centre, NO-9296 Tromsø, Norway. 15 16 * Corresponding author: 17 Andrea Balboni 18 Department of Veterinary Medical Sciences, Alma Mater Studiorum-University of Bologna, Via Tolara di Sopra 50, 19 40064, Ozzano dell’Emilia (BO), Italy. 20 Tel. +39 051 2097083 21 e-mail: a.balboni@unibo.it 22 ORCID: orcid.org/0000-0002-8049-6645 23 24 Acknowledgements 25 We thank all the trappers and hunters that provided the fox samples. Funding for sampling the fox tissues were provided 26 from the Norwegian Polar Institute and financed by the Norwegian Environment Agency to Climate –ecological 27 Observatory for Arctic Tundra (COAT). We also acknowledge Dr. Javier Sánchez Romano for help with the tissue 28 samples and DNA extraction. 29