Vol.:(0123456789) 1 3
Archives of Microbiology
https://doi.org/10.1007/s00203-019-01626-y
ORIGINALPAPER
Impact of seasonality and environmental conditions on yeast diversity
from camel’s milk collected in Algeria
Soufan Ider
1,2,3
· Yanath Belguesmia
2
· Françoise Coucheney
2
· Mabrouk Kihal
1
· Djamel Drider
2
Received: 2 January 2019 / Revised: 22 January 2019 / Accepted: 30 January 2019
© Springer-Verlag GmbH Germany, part of Springer Nature 2019
Abstract
During this study, we characterized the seasonality’s impact and environmental conditions on the yeast diversity from raw
camel’s milk collected in Algeria. The yeast counts were estimated to 3.55 × 10
2
CFU mL
−1
, with a maximum of 6.3 × 10
2
CFU mL
−1
. The yeasts were categorized phenotypically by API 20C AUX, MALDI-TOF and genetically by sequencing
26S rDNA and ITS1-5.8S-ITS2. The rDNA sequencing approaches revealed 12 species including unusual ones such as
Trichosporon asahii, Pichia fermentans, Millerozyma farinosa, Pichia galeiformis, Candida tartarivorans and Pichia man-
shurica. The most dominant species were T. asahii (23%), P. fermentans (19%) and Rhodotorula mucilaginosa (14%). The
high occurrence and large diversity were registered in samples collected during the autumn season, in the semi-arid and arid
highlands regions with 0.66 × 10
3
CFU mL
−1
and 0.51 × 10
3
CFU mL
−1
, respectively. Interestingly, T. asahii, R. mucilaginosa,
P. fermentans, C. parapsilosis and C. zeylanoides were detected during both spring and autumn.
Keywords Camel milk · Yeasts · Biodiversity · Environmental conditions · Phenotypic and genotypic characterization
Introduction
The Camelus dromedarius is a single-humped camel spe-
cies belonging to the Camelidae family (Siboukeur 2007). In
Algeria, national camel reserves were estimated, more than
a decade ago to 150 thousand animals, and this trend was
accompanied by an increase in the number of camel milk
producers (Lhoste 2004). The geographical distribution of
the camel population in Algeria encompasses the semi-arid
areas, the arid areas and the full desert. Camels living in
these areas are subjected to highly elevated temperatures
and limited access to water sources. In spite of these harsh
environmental conditions, this animal produces milk almost
yearlong and in amounts considered to be greater than those
from other animals living in same conditions (Park and
Haenlein 2008).
Camel’s milk is rich in proteins, fats, minerals and vita-
mins (Konuspayeva et al. 2009; Haddadin et al. 2008).
Additionally, camel’s milk is characterized for its high
water level, near-neutral pH and high lactose (http://www.
fao.org/docre p/003/X6528 E/X6528 E03.htm). This milk
content can permit the development of a rich autochtho-
nous microbiota. Microorganisms present in such ecosys-
tem are basically issued from teat apex, milking equip-
ment, air, water, food, grass, soil and other environments
(Coorevits et al. 2008; Angulo et al. 2009; Vacheyrou et al.
2011). Yeasts were found in the raw milk and their occur-
rence was estimated to be about 10
4
CFU mL
−1
, mostly
belonging to Kluyveromyces lactis, Zygosaccharomyces
bailii, Zygosaccharomyces bisporus and Debaryomyces
hansenii species with no pathogenic yeasts identifed in
camel milk (Baroiller and Schmidt 1990; Njage et al.
2011; Yam et al. 2015). Their diversity and amount in the
raw milk was reported to be rearing practices dependent.
In fact, yeasts with capabilities to metabolize lactose, con-
versely to those devoid of such metabolic activity, can eas-
ily grow and take advantage from media containing lactose
Communicated by Erko Stackebrandt.
* Djamel Drider
djamel.drider@univ-lille.fr
1
Laboratoire de Microbiologie Fondamentale et Appliquée,
Faculté des Sciences de la Nature et de la Vie, Université
Oran 1 - Ahmed Ben Bella, Es Senia, 31100 Oran, Algeria
2
EA 7394-Institut Charles Violette, Université de Lille,
59000 Lille, EA, France
3
Present Address: Département de Biologie, Faculté des
Sciences de la Nature et de la Vie et Sciences de la Terre,
Université de Ghardaïa, BP 455, 47000 Ghardaïa, Algeria