This article is protected by copyright. All rights reserved RESEARCH ARTICLE Protective effects of camel whey protein against scrotal heat-mediated damage and infertility in the mouse testis through YAP/Nrf2 and PPAR-gamma signaling pathways † Running title: Camel whey protein improves testis injury induced by heat stress Gamal Badr 1, 2* , Hanem S. Abdel-Tawab 1 , Nancy K. Ramadan 2, 3 , Samia F. Ahmed 3 , and Mohamed H. Mahmoud 4,5 1 Zoology Department, Faculty of Science, Assiut University, 71516 Assiut, Egypt 2 Laboratory of Immunology & Molecular Physiology, Zoology Department, Faculty of Science, Assiut University, 71516 Assiut, Egypt 3 Animal Health Research Institute, Assiut Branch, 71526 Assiut, Egypt 4 Deanship of Scientific Research, King Saud University, 11692 Riyadh, Saudi Arabia 5 Food Science and Nutrition Department, National Research Center, Dokki, 12622 Cairo, Egypt † This article has been accepted for publication and undergone full peer review but has not been through the copyediting, typesetting, pagination and proofreading process, which may lead to differences between this version and the Version of Record. Please cite this article as doi: [10.1002/mrd.22987] Received 5 January 2018; Revised 7 April 2018; Accepted 16 April 2018 Molecular Reproduction & Development This article is protected by copyright. All rights reserved DOI 10.1002/mrd.22987