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*Corresponding author.
Email: wibowo.mangun@yahoo.co.id / wibowo.mangun@ui.ac.id
International Food Research Journal 21(2): 527-532 (2014)
Journal homepage: http://www.ifrj.upm.edu.my
1
Sulistiani,
2
Abinawanto,
3
Endang Sukara,
2
Andi Salamah,
4
Achmad Dinoto and
2*
Wibowo Mangunwardoyo
1
Graduate Student, Department Biology, Faculty of Mathematic and Natural Sciences, Universitas Indonesia,
Depok, Indonesia, 16424
2
Department Biology, Faculty of Mathematic and Natural Sciences, Universitas Indonesia, Depok, Indonesia,
16424
3
Research Center for Biotechnology, Indonesian Institute of Sciences (LIPI), Jl. Raya Jakarta Bogor Km 46
Cibinong, West Java, Indonesia, 16911
4
Research Center for Biology, Indonesian Institute of Sciences (LIPI), Jl. Raya Jakarta Bogor Km 46
Cibinong, West Java, Indonesia, 16911
Identifcation of lactic acid bacteria in sayur asin from Central Java
(Indonesia) based on 16S rDNA sequence
Abstract
Sayur asin is commonly known in Indonesia, in particular Central Java, as a spontaneous
fermentated mustard product by epiphytic lactic acid bacteria (LAB) from the plants. However,
information regarding LAB diversity in Indonesia is lacking and fragmentary. Therefore,
efforts on inventory of LAB from sayur asin in Indonesia has been conducted since April 2013
in order to determine its diversity. In the current study, LAB diversity in fve samples of sayur
asin collected from Solo, Yogyakarta, and Semarang (Central Java) is reported. LAB were
isolated from both of fermented mustards and fermenting liquors. Molecular identifcation of
the isolates was conducted based on 16S rDNA sequence data. A total of 246 isolates of LAB
was successfully isolated and identifed. The bacteria belong to 11 species, viz, Lactobacillus
farciminis (15 isolates), L. fermentum (83 isolates), L. namurensis (18 isolates), L. plantarum
(107 isolates), L. helveticus (1 isolate), L. brevis (1 isolate), L. versmoldensis (3 isolates), L.
casei (12 isolates), L. rhamnosus (2 isolates), L. fabifermentans (3 isolates) and L. satsumensis
(1 isolate). The current study revealed that L. plantarum and L. fermentum as common LAB
used in sayur asin production from Central Java, Indonesia.
Introduction
Sayur asin has been known as traditional
fermented mustard produced and consumed in many
areas in Indonesia for years (Puspito and Graham,
1985). Manufacture of fermented mustard was
made by addition of the salt to vegetables, allowing
the growth of certain fermentable microorganisms,
resulting in sensory changes bearing acidic and unique
charateristics to the mustard (Chiou, 2004). Epiphytic
lactic acid bacteria (LAB) of which initially amounted
to slightly between 10-1000 cfu/g plant (0.001 to 1%
of the total population of microorganisms) became
dominant within the microorganism population in in
the fermented mustard as a result of the addition on
the anaerobic treatment and salt on the vegetables
(Daeschel et al., 1987; Azcarate and Todd, 2010).
According to Chao et al. (2009), fermented
mustard is a good source for the exploration of
LAB. Differences in the manufacture of fermented
mustard, salt concentration, temperature during the
fermentation process, and the level of anaerobiosis
affect the selection of certain LAB species of in the
mustard fermentation. Puspito and Graham (1985)
reported that there was a difference in manufacturing
techniques in the mustard fermentation in Indonesia.
Several locations in central Java, such as Solo,
Yogyakarta and Semarang, have traditionally been
known as producers of sayur asin in Indonesia.
Therefore, exploration of LAB in these areas will
be important to reveal LAB diversity in Indonesian
traditional fermented mustard.
In the development of LAB taxonomy,
several species of LAB heve been recognized as
complex species, which is diffcult to distinguish
by conventional techniques, such as physiological
and biochemical tests. Therefore, increasing the
numbers of LAB species which have phenotypic
and physiological similarities using the conventional
techniques often resulting in the misidentifcation
(Vogel and Matthias, 2008; Nikolova et al., 2009).
In addition, the conventional techniques are also
time-consuming, not reproducible, and do not give
accurate results (Yu et al., 2009). Development of
molecular methods based on nucleotide sequence
data from ribosomal DNA has complemented
Keywords
Diversity
Fermented mustard
Lactobacillus
16S rDNA
Indonesia
Article history
Received: 1 November 2013
Received in revised form:
6 December 2013
Accepted: 12 December 2013