Pharmacology Biochemistry & Behavior, Vol. 15, pp. 513-514, 1981, Printed in the U.S.A.
Naloxone Potentiates Shock-Induced
Aggressive Behavior in Mice
STEFANO PUGLISI-ALLEGRA AND ALBERTO OLIVERIO
lstituto di Psicobioiogia e Psicofarmacologia, via Reno, 1 - 00198 Roma, Italy and
Istituto di Fisiologia Generale, Universit~ di Roma
Received 19 February 1981
PUGLISI-ALLEGRA, S. AND A. OLIVERIO. Naloxone potentiates shock-induced aggressive behavior in mice.
PHARMAC. BIOCHEM. BEHAV. 15(3) 513-514, 1981.--Naloxone (0.025 to 0.05 mg/kg IP) potentiates shock-induced
aggressive behavior in C57BL/6 mice but not in DBA/2 mice which do not fight in absence of drug, This effect is not related
to pain sensitivity since the doses used in this experiment do not lower tail-flick threshold in C57BL/6 mice. These f'mdings
are discussed in terms of the role of the endorphin system and of catecholamine related sensory attention in a number of
social interactions in the mouse.
Naloxone Shock-induced aggressive behavior Endorphins Inbred mice
ENDOGENOUS opioids (enkephalins and endorphins) are
involved in a number of behavioral patterns and social activ-
ities connected to emotionality. Animals treated with nalox-
one, a drug which antagonizes the effects of endorphins, are
characterized by increased locomotor activity, different
signs of social discomfort and enhanced sexual behavior (for
review see Panksepp et al. [5]).
These behavioral patterns were described in different
species ranging from birds to rodents and the functions of the
endogenous analgesic systems have been interpreted in
terms of adaptive species-specific mechanisms. Differences
in the function of the endorphin system and in the behavioral
effects of opiates have been described and interpreted in
terms of different types of receptors and of their cerebral
distribution [9]. In particular morphine or other opiates exert
a stimulating effect on the activity of some strains of mice
such as C57BL/6 (C57), while they depress or induce freez-
ing responses in other strains such as DBA/2 (DBA); these
differences have been explained in terms of endogenous
opiate receptors and of dopaminergic and noradrenergic re-
sponses [2]. C57 and DBA mice are also characterized by
different levels of aggressive behavior in a number of tests in
response to pharmacological and environmental manipula-
tions [6,10]. In the present study these strains of mice were
injected with naloxone in order to assess whether or not an
inhibition of endogenous opiates results in an increased
shock-induced aggressive behavior.
METHOD
Male C57BL/6 (C57) and DBA/2 (DBA) mice (Charles
River, Como, Italy), 11-12 weeks old were used. Shock-
induced aggressive behavior was measured in an apparatus
in which the bites between two mice are recorded through a
counter [7]. The apparatus consists of an experimental
chamber (9x9x14 cm). Two holes (1 cm diameter at 1 cm
above the grid floor) allow the tails to extend out of the
chamber on two opposite walls. A threaded bush previously
glued on each tail by means of Bostik is fixed by a nut in the
center of a ball bearing, so that the mouse can move freely
with no danger to the tail. The two ball bearings work as slip
rings for a contact detector. A low current (2 t~A) passing
through both mice during biting is amplified by the contact
detector so that all bites, but no other body contact, can be
recorded by a digital counter. Aggressive behavior was in-
duced by shocking simultaneously both mice. The electric
shock (0.25 mA) was applied by means of two electrodes to
the part of the tail available between the hole and the ball
bearing.
In our experiments each treatment group consisted of ten
pairs of mice per strain. Each session lasted for four minutes
in which shocks of 0.5 sec duration were delivered every 3
sec. Prior to each encounter the mice were placed in oppo-
site halves of the experimental cage, separated by a sliding
door. After a 60 sec adaptation period the sliding door was
raised, and shocks were delivered. The experimental cage
was cleaned between sessions in order to remove odors.
Mice of each pair came from different breeding groups. To
study pain sensitivity the tail-flick method was used.
Naloxone hydrochloride was injected intraperitoneally 10
minutes before testing at different doses. Control mice were
injected with saline.
Data were analyzed statistically by single-factor analysis
of variance (ANOVA; independent). Further analysis for
individual between-group comparisons was carried out by
employing the error term of the overall analysis of variance.
RESULTS
Saline-injected C57 mice are characterized by very low
levels of aggressive responses: Figure I shows that naloxone
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