1 Research Article Activities of carbohydrate-metabolism enzymes in pre-drought primed wheat plants under drought stress during grain filling Running title: Carbohydrate metabolism in drought primed wheat plants Muhammad Abid 1, 2† , Zhongwei Tian 1† , Jinling Hu 1 , Attiq Ullah 1 , Yakun Cui 1, 3 , Shafaqat Ali 4 , Suyu Jiang 1 , Rizwan Zahoor 1 , Yonghui Fan 1 , Jiang Dong 1 , Tingbo Dai 1** 1 . Key Laboratory of Crop Physiology, Ecology and Production Management, Nanjing Agricultural University, Nanjing 210095, China 2 . Department of Soil Conservation, Narowal 51600, Pakistan 3 . Institute of Food Crops, Jiangsu Academy of Agricultural Sciences, Nanjing 210014, China 4 . Department of Environmental Sciences and Engineering, Government College University, Allama Iqbal Road 38000, Faisalabad, Pakistan † These authors contributed equally to this work * Correspondence: Tingbo Dai (tingbod@njau.edu.cn) Edited by: Steven M. Smith, University of Tasmania, Australia This article has been accepted for publication and undergone full peer review but has not been through the copyediting, typesetting, pagination and proofreading process, which may lead to differences between this version and the Version of Record. Please cite this article as doi: [10.1111/jipb.12628] This article is protected by copyright. All rights reserved. Received: September 12, 2017; Accepted: December 17, 2017